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Selected articles

Tyrannosaurus is a famous dinosaur genus and a fixture in popular culture. Known colloquially as T. rex, the species Tyrannosaurus rex hails from what is now western North America. Some scientists consider the slightly older Tarbosaurus bataar from Asia to represent a second species of Tyrannosaurus, while others maintain Tarbosaurus as a separate genus. Like other tyrannosaurid theropods, Tyrannosaurus was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to the large and powerful hindlimbs, Tyrannosaurus forelimbs were small, and retained only two digits. Although other theropods rivaled or exceeded T. rex in size, it was the largest known tyrannosaurid and one of the largest known land predators, measuring over 12 meters (40 feet) in length and weighing as much as an elephant. Fossils of T. rex have been found in North American rock formations dating to the very end of the Cretaceous Period; it was among the last dinosaurs to exist prior to the Cretaceous-Tertiary extinction event. More than 30 specimens of T. rex have now been identified, some nearly complete, which has allowed significant research into many aspects of its biology, including its life history and biomechanics. However, the feeding habits and potential speed of T. rex remain controversial. (see more...)

Iguanodon is a genus of ornithopod dinosaur that lived roughly halfway between the first of the swift bipedal hypsilophodontids and the ornithopods' culmination in the duck-billed dinosaurs. Many species of Iguanodon have been named, dating from the Kimmeridgian age of the Late Jurassic Period to the Cenomanian age of the Late Cretaceous Period from Asia, Europe, and North America. However, research in the first decade of the 21st century suggests that there is only one well-substantiated species named I. bernissartensis, that lived from the late Barremian to the earliest Aptian ages (Early Cretaceous) in Belgium, between about 126 and 125 million years ago. Iguanodon's most distinctive features were its large thumb spikes, which were possibly used for defence against predators, combined with long prehensile fifth fingers able to forage for food. Named in 1825 by English geologist Gideon Mantell, Iguanodon was the second dinosaur formally named, after Megalosaurus. A large, bulky herbivore, Iguanodon is a member of Iguanodontia, along with the duck-billed hadrosaurs. (see more...)

Triceratops was a herbivorous genus of ceratopsid dinosaur that lived during the late Maastrichtian around 68-65 million years ago in what is now North America; it was one of the last dinosaurs to appear before the great K-T extinction event. Bearing a large bony frill and three horns on its large four-legged body, and conjuring similarities with the modern rhinoceros, Triceratops is one of the most recognizable of all dinosaurs. Though it shared the landscape with, and was preyed upon by, the fearsome Tyrannosaurus, it is unclear whether the two battled the way they are commonly depicted in movies and children's dinosaur books. Although no complete skeleton has been found, Triceratops is well known from numerous partial specimens collected since the genus' introduction in 1887. The function of their frills and three distinctive facial horns has long inspired debate. Although traditionally viewed as defensive weapons against predators, the latest theories suggest these features were primarily used in display for courtship and dominance, much like the antlers and horns of modern reindeer, mountain goats or rhinoceros beetles. (see more...)

Modern depiction of Diplodocus carnegiei.

Diplodocus (meaning 'double bar') is a genus of diplodocid sauropod dinosaur whose fossilised skeleton was first discovered in 1878. The generic name refers to its double-beamed chevron bones (Greek diplos/διπλος meaning 'double' and dokos/δοκος meaning 'wooden beam' or 'bar') located in the underside of the tail. They were initially believed to be unique to Diplodocus; however, they have since then been discovered in other diplodocids.

It lived in what is now western North America at the end of the Jurassic Period. Diplodocus was one of the more common dinosaurs found in the Upper Morrison Formation, about 150 to 147 million years ago, in what is now termed the Kimmeridgian and Tithonian stages. This was an environment and time dominated by gigantic sauropod dinosaurs such as Camarasaurus, Barosaurus, Apatosaurus and Brachiosaurus. Diplodocus is among the most easily identifiable dinosaurs, with its classic dinosaur shape, long neck and tail and four sturdy legs. For many years, it was the longest dinosaur known. Its great size may have been a deterrent to the predators Allosaurus and Ceratosaurus: their remains have been found in the same strata, which suggests they coexisted with Diplodocus. (see more...)

Stegosaurus skeleton as drawn by Othniel Charles Marsh

Stegosaurus (meaning 'roof-lizard') is a genus of stegosaurid armoured dinosaur from the Late Jurassic period (Kimmeridgian to Early Tithonian) in what is now western North America. However, in 2006 a specimen of Stegosaurus was announced from Portugal, suggesting that they were present in Europe as well. Due to its distinctive tail spikes and plates, Stegosaurus is one of the most recognisable dinosaurs, along with Tyrannosaurus, Triceratops and Apatosaurus. At least three species have been identified in the upper Morrison Formation and are known from the remains of about 80 individuals. They lived some 155 to 145 million years ago, in an environment and time dominated by the giant sauropods Diplodocus, Camarasaurus and Apatosaurus. A large, heavily-built and herbivorous quadruped, Stegosaurus had a distinctive and unusual posture, with a heavily-arched back, short forelimbs, head held low to the ground and a stiffened tail held high in the air. Its array of plates and spikes have been the subject of much speculation. The spikes were most likely used for defence, while the plates have also been proposed as a defensive mechanism, as well as having display and thermoregulatory (heat control) functions. Stegosaurus was the largest of all the stegosaurians (bigger than genera such as Kentrosaurus and Huayangosaurus) and, although roughly bus-sized, it nonetheless shared many anatomical features (including the tail spines and plates) with the other stegosaurian genera. (see more...)

Velociraptor (meaning 'swift thief') was a genus of dromaeosaurid theropod dinosaur that existed approximately 83 to 70 million years ago during the later part of the Cretaceous Period. There is only one currently recognized species, V. mongoliensis, although others have been assigned in the past. Fossils of this species have been found in central Asia, from both Inner and Outer Mongolia.

Smaller than other dromaeosaurids like Deinonychus and Achillobator, the turkey-sized Velociraptor nevertheless shared many of the same anatomical features. It was a bipedal carnivore with a long, stiffened tail and had an enlarged, sickle-shaped claw on each hindfoot, which is thought to have been used to kill its prey. Velociraptor can be distinguished from other dromaeosaurids by its long and low skull, with an upturned snout.

Due in large part to its prominent role in Michael Crichton's novel Jurassic Park and the subsequent motion picture series, Velociraptor (commonly shortened to 'raptor') is one of the dinosaur genera most familiar to the general public. It is also well-known to paleontologists, with over a dozen recovered fossil skeletons — the most of any dromaeosaurid. One particularly famous specimen shows a Velociraptor locked in combat with a Protoceratops. (see more...)

Compsognathus is a monotypic genus of small, bipedal, carnivorous theropod dinosaur. The species Compsognathus longipes was the size of a turkey and lived around 150 million years ago, the early Tithonian stage of the late Jurassic Period, in what is now Europe. Paleontologists have found two well-preserved fossils, one in Germany in the 1850s and the second in France more than a century later. Many presentations still describe Compsognathus as a "chicken-sized" dinosaur because of the small size of the German specimen, which is now believed to be a juvenile form of the larger French specimen. Compsognathus is one of the few dinosaurs for which the diet is known with certainty: the remains of small, agile lizards are preserved in the bellies of both specimens. Although not recognized as such at the time of its discovery, Compsognathus is the first theropod dinosaur known from a reasonably complete fossil skeleton. Until the 1990s, it was the smallest known non-avialan dinosaur and the closest supposed relative of the early bird Archaeopteryx. Thus, its genus is one of the few dinosaur genera to be well known outside of paleontological circles. (see more...)

Albertosaurus (meaning 'Alberta lizard') was a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period, more than 70 million years ago. The type species, A. sarcophagus, was restricted in range to the modern-day Canadian province of Alberta, after which the genus is named. Scientists disagree on the number of species represented in the genus, recognizing either one or two species. As a tyrannosaurid, Albertosaurus was a bipedal predator with a massive head, jaws lined with dozens of large teeth and tiny, two-fingered 'hands' and it may have been at the top of the food chain in its local ecosystem. Although relatively large for a theropod, Albertosaurus was much smaller than its more famous relative Tyrannosaurus, probably weighing only as much as a modern black rhinoceros. Fossils of more than twenty individuals have been recovered, providing scientists with a more detailed knowledge of Albertosaurus anatomy than is available for other tyrannosaurids. The discovery of ten individuals at one site provides evidence of pack behavior and allows studies of developmental biology which are impossible with lesser-known animals. (see more...)

Psittacosaurus is a genus of psittacosaurid ceratopsian dinosaur from the Early Cretaceous Period of what is now Asia, about 130 to 100 million years ago. It is notable for being the most species-rich dinosaur genus. Nine to eleven species are recognized from fossils found in different regions of modern-day China, Mongolia and Russia, with a possible additional species from Thailand. All species of Psittacosaurus were gazelle-sized bipedal herbivores characterized by a high, powerful beak on the upper jaw. At least one species had long, quill-like structures on its tail and lower back, possibly serving a display function. Psittacosaurus is not as familiar to the general public as its distant relative Triceratops but it is one of the most completely known dinosaur genera. Fossils of over 400 individuals have been collected so far, including many complete skeletons. Most different age classes are represented, from hatchling through to adult, which has allowed several detailed studies of Psittacosaurus growth rates and reproductive biology. The abundance of this dinosaur in the fossil record has led to establishing the Psittacosaurus biochron for the Early Cretaceous of east Asia. (see more...)

Archaeopteryx is the earliest and most primitive bird known to date. It lived in the late Jurassic Period around 155-150 million years ago in what is now southern Germany. At the time Archaeopteryx lived, Europe was an archipelago of islands in a shallow warm tropical sea, much closer to the equator than it is now. Archaeopteryx had feathers and wings, but it also had teeth and a skeleton similar to a small carnivorous dinosaur; therefore, it had both bird and theropod dinosaur features. Similar in size and shape to a European Magpie, it bore broad, rounded wings and a long tail. Archaeopteryx could grow to about half a metre, or 1.6 feet in length. Its feathers resembled the flight feathers of modern birds, suggesting not only capacity for flight, but also homoiothermy. Otherwise, its features were reptilian, with jaws lined with sharp teeth, three 'fingers' ending in curved claws and a long bony tail. These features, which are consistent with theropod dinosaurs, made Archaeopteryx a hot topic in the debate on evolution. Many have seen it as a true 'missing link'. In 1862 the description of the first intact specimen of Archaeopteryx, just two years after Charles Darwin published The Origin of Species, set off a firestorm of debate about evolution and the role of transitional fossils that endures to this day. The eleven fossils currently classified as Archaeopteryx are the oldest evidence of feathers on the planet and the only ones dated from Jurassic times. (see more...)

Thescelosaurus neglectus drawing.

Thescelosaurus is a genus of small ornithopod dinosaur known from Upper Cretaceous rocks of western North America. It belonged to the last dinosaurian fauna of North America before the Cretaceous–Paleogene extinction event at approximately 66 million years ago, living alongside dinosaurs such as Tyrannosaurus and Triceratops. This common genus was described from a specimen discovered in 1891, but not unpacked and studied until the 1910s. These circumstances suggested the names of the genus and type species T. neglectus, which roughly translate to "godlike, wondrous, or marvelous neglected lizard". Thescelosaurus is best known from several partial skeletons representing three species: T. neglectus, T. assiniboiensis, and T. garbanii. One specimen unearthed in 1993 was initially thought to include a preserved heart, but later study found the object is probably a concretion. Thescelosaurus was a bipedal animal with a relatively long pointed skull and robust limbs. Typical individuals measured on the order of 2.5 to 4.0 metres (8.2 to 13.1 ft) long. It was probably primarily herbivorous and may have preferred to live near streams. (see more...)

Styracosaurus was a genus of herbivorous ceratopsian dinosaur from the Cretaceous Period (Campanian stage), about 76.5 to 75.0 million years ago. It had four to six long horns extending from its neck frill, a smaller horn on each of its cheeks, and a single horn protruding from its nose, which may have reached dimensions of around 60 centimeters (2 ft) long and 15 centimeters (6 in) wide. The function or functions of the horns and frills have been the subject of debate for many years. Styracosaurus was a large dinosaur, reaching lengths of 5.5 meters (18 ft) and weighing nearly 3 tons. It stood about 1.8 meters (6 ft) tall. Styracosaurus possessed four short legs and a bulky body. Its tail was rather short. It also had a beak and flat cheek teeth, indicating that its diet was herbivorous. Like other ceratopsians, this dinosaur may have been a herd animal, traveling in large groups, as suggested by bonebeds. Named by Lawrence Lambe in 1913, Styracosaurus is a member of the Centrosaurinae. Two species, S. albertensis and S. ovatus are currently assigned to Styracosaurus. Other species assigned to the genus have since been reassigned elsewhere. (see more...)

Daspletosaurus (meaning 'frightful lizard') was a genus of tyrannosaurid theropod dinosaur that lived in western North America between 80 and 73 million years ago, during the Late Cretaceous Period. Fossils of the only named species (D. torosus) were found in Alberta, although other possible species from Alberta, Montana and New Mexico await description. Including these undescribed species, Daspletosaurus is the most species-rich genus of tyrannosaur.

Daspletosaurus is closely related to the much larger and more recent Tyrannosaurus. Like most known tyrannosaurids, it was a multi-ton bipedal predator equipped with dozens of large, sharp teeth. Daspletosaurus had the small forelimbs typical of tyrannosaurids, although they were proportionately longer than in other genera. It was probably similar in weight to a modern white rhinoceros or a small elephant.

As an apex predator, Daspletosaurus was at the top of the food chain, probably preying on large dinosaurs like the ceratopsid Centrosaurus and the hadrosaur Hypacrosaurus. In some areas, Daspletosaurus coexisted with another tyrannosaurid, Gorgosaurus, though there is some evidence of niche differentiation between the two. While Daspletosaurus fossils are rarer than other tyrannosaurids, the available specimens allow some analysis of the biology of these animals, including social behavior, diet and life history. (see more...)

Deinonychus (meaning 'terrible claw') was a genus of carnivorous dromaeosaurid dinosaur. This 3-4 metre (10-13 ft) long dinosaur lived during the early Cretaceous Period (Aptian - Albian stages, 121 to 98.9 million years ago). Fossils of the only named species (D. antirrhopus) have been recovered from Montana, Wyoming and Oklahoma, though teeth that may belong to Deinonychus have been found much farther east in Maryland.

Its name refers to the unusually large, sickle-shaped talon on the second toe of each hind foot, which was probably held retracted while the dinosaur walked on the third and fourth toes. It was commonly thought that Deinonychus would kick with the sickle claw to slash at its prey but recent tests on reconstructions of similar Velociraptor talons suggest that the claw was used to stab, not slash. As in other dromaeosaurids, the tail was stiffened by a series of elongated bones and bone processes. This might have given Deinonychus greater balance and turning ability. In both the Cloverly and Antlers Formation, Deinonychus remains have been found closely associated with those of the ornithopod Tenontosaurus. Teeth discovered associated with Tenontosaurus specimens imply it was hunted or at least scavenged upon by Deinonychus.

Paleontologist John Ostrom's study of Deinonychus in the late 1960s revolutionized the way scientists thought about dinosaurs, igniting the debate on whether or not dinosaurs were warm-blooded. Before this, the popular conception of dinosaurs had been one of plodding, reptilian giants. Ostrom noted lightweight bones and stiffened tendons which revealed an active, agile predator. (see more...)

Parasaurolophus (meaning 'near crested lizard') was a genus of ornithopod dinosaur from the Late Cretaceous Period of what is now North America, about 76-73 million years ago. It was a herbivore that walked both as a biped and a quadruped. Three species are recognized: P. walkeri (the type species), P. tubicen, and the short-crested P. cyrtocristatus. Remains are known from Alberta (Canada), and New Mexico and Utah (United States). It was first described in 1922 by William Parks from a skull and partial skeleton in Alberta. Parasaurolophus is a hadrosaur, part of a diverse family of Cretaceous dinosaurs known for their range of bizarre head adornments. This genus is known for its large, elaborate cranial crest, which at its largest forms a long curved tube projecting upwards and back from the skull. A similar crest is found on Charonosaurus from China, which may have been its closest relative. The crest has been much discussed by scientists; the consensus is that major functions included visual recognition of both species and gender, acoustic resonance, and thermoregulation. It is one of the rarer duckbills, known from only a handful of good specimens. (see more...)

Majungasaurus (meaning 'Mahajanga lizard') is a genus of abelisaurid theropod dinosaur that lived in Madagascar from 70 to 65 million years ago, at the end of the Cretaceous Period. Only one species (M. crenatissimus) has been identified. This dinosaur was briefly called Majungatholus, a name which is now considered a junior synonym of Majungasaurus.

Like other abelisaurids, Majungasaurus was a bipedal predator with a short snout. Although the forelimbs are not completely known, they were very short, while the hindlimbs were longer and very stocky. It can be distinguished from other abelisaurids by its wider skull, the very rough texture and thickened bone on the top of its snout, and the single rounded horn on the roof of its skull, which was originally mistaken for the dome of a pachycephalosaur. It also had more teeth in both upper and lower jaws than most abelisaurids.

Known from several well-preserved skulls and abundant skeletal material, Majungasaurus has recently become one of the best-studied theropod dinosaurs from the Southern Hemisphere. It appears to be most closely related to abelisaurids from India rather than South America or continental Africa, a fact which has important biogeographical implications. Majungasaurus was the apex predator in its ecosystem, mainly preying on sauropods like Rapetosaurus, and is also the only dinosaur for which direct evidence of cannibalism is known. (see more...)

The Cretaceous–Paleogene extinction event, also known as either the K–Pg extinction event, Cretaceous–Tertiary extinction event, or K–T extinction event, was the large-scale mass extinction of animal and plant species in a geologically short period of time, approximately 66 million years ago (mya). The Cretaceous–Paleogene extinction event is associated with a geological signature, usually a thin band dated to that time and found in various parts of the world, known as the K–Pg boundary. K is the traditional abbreviation for the Cretaceous Period, Pg is the abbreviation for the Paleogene Epoch, and T is the abbreviation for the Tertiary Period. The event marks the end of the Mesozoic Era, and the beginning of the Cenozoic Era.

Non-avian dinosaur fossils are only found below the K–Pg boundary and became extinct immediately before or during the event. A very small number of dinosaur fossils have been found above the K-Pg boundary, but they have been explained as reworked, that is, fossils that have been eroded from their original locations then preserved in later sedimentary layers. Mosasaurs, plesiosaurs, pterosaurs and many species of plants and invertebrates also became extinct. Mammalian and bird clades passed through the boundary with few extinctions, and radiation from those Maastrichtian clades occurred well past the boundary. Rates of extinction and radiation varied across different clades of organisms.

Many scientists theorize that the K-Pg extinctions were caused by one or more catastrophic geological events such as massive asteroid impacts or increased volcanic activity. Several impact craters and massive volcanic activity in the Deccan traps have been dated to the approximate time of the extinction event. These geological events may have reduced sunlight and hindered photosynthesis, leading to a massive disruption in Earth's ecology. Other researchers believe the extinction was more gradual, resulting from slower changes in sea level or climate. (see more...)

Lambeosaurus life restoration

Lambeosaurus (meaning 'Lambe's lizard') is a genus of hadrosaurid dinosaur that lived about 76 to 75 million years ago, in the Late Cretaceous Period (Campanian) of North America. This bipedal/quadrupedal, herbivorous dinosaur is known for its distinctive hollow cranial crest, which in the best-known species resembled a hatchet. Several possible species have been named, from Alberta (Canada), Montana (United States), and Baja California (Mexico), but only the two Canadian species are currently well known. At about 15 meters (50 ft) long, the Mexican species L. laticaudus was one of the longest ornithischians. The other species were more modestly sized. Lambeosaurus was belatedly described in 1923 by William Parks, over twenty years after the first material was studied by Lawrence Lambe. The genus has had a complicated taxonomic history, in part because small-bodied crested hadrosaurids now recognized as juveniles were once thought to belong to their own genera and species. Currently, the various skulls assigned to the type species L. lambei are interpreted as showing age differences and sexual dimorphism. Lambeosaurus was closely related to the better known Corythosaurus, which is found in slightly older rocks, as well as the less well-known genera Hypacrosaurus and Olorotitan. All had unusual crests, which are now generally assumed to have served social functions like noisemaking and recognition. (see more...)

Acrocanthosaurus (meaning 'high-spined lizard') was a genus of allosauroid theropod dinosaur that existed in what is now North America during the mid-Cretaceous Period, approximately 125 to 100 million years ago. Like most dinosaur genera, Acrocanthosaurus contains only a single species, A. atokensis. Its fossil remains are found mainly in the U.S. states of Oklahoma and Texas, although teeth attributed to Acrocanthosaurus have been found as far east as Maryland.

Acrocanthosaurus was a bipedal predator. As the name suggests, it is best known for the high neural spines on many of its vertebrae, which most likely supported a ridge of muscle over the animal's neck, back and hips. Acrocanthosaurus was one of the largest theropods, approaching 12 meters (40 ft) in length, and weighing up to about 2.40 metric tons (2.65 short tons). Large theropod footprints discovered in Texas may have been made by Acrocanthosaurus, although there is no direct association with skeletal remains.

Recent discoveries have elucidated many details of its anatomy, allowing for specialized studies focusing on its brain structure and forelimb function. However, there is still debate over its evolutionary relationships, with some scientists classifying it as an allosaurid, and others as a carcharodontosaurid. Acrocanthosaurus was the largest theropod in its ecosystem and likely an apex predator which possibly preyed on large sauropods and ornithopods. (see more...)

Chicxulub Crater is an ancient impact crater buried underneath the Yucatán Peninsula, with its center located near the town of Chicxulub, Yucatán, Mexico. The crater is over 180 kilometers (110 mi) in diameter, making the feature one of the largest confirmed impact structures in the world; the asteroid or comet whose impact formed the crater was at least 10 km (6 mi) in diameter. The crater was named for the nearby town, as well as for the literal Maya translation of the name: "tail of the devil." The crater was discovered by Glen Penfield, a geophysicist who had been working in the Yucatán while looking for oil during the late 1970s. The presence of tektites, shocked quartz and gravity anomalies, as well as the age of the rocks and isotope analysis, show that this impact structure dates from the late Cretaceous Period, roughly 65 million years ago. The impact associated with the crater is implicated in causing the extinction of the dinosaurs as suggested by the Cretaceous–Paleogene boundary, although some critics disagree that the impact was the sole reason and also debate whether there was a single impact or whether the Chicxulub impactor was one of several that may have struck the Earth at around the same time. Recent evidence suggests that the impactor was a piece of a much larger asteroid which broke up in a collision more than 160 million years ago.(see more...)

Allosaurus (meaning 'strange lizard') is a genus of large theropod dinosaur that lived 155 to 145 million years ago, in the late Jurassic period. The first remains that can definitely be ascribed to this genus were described in 1877 by Othniel Charles Marsh. As one of the first well-known theropod dinosaurs, it has long attracted attention outside of paleontological circles, and has been a lead dinosaur in several films and documentaries.

Allosaurus was a large bipedal predator with a large skull, equipped with dozens of large, sharp teeth. It averaged 8.5 meters (30 ft) in length, though fragmentary remains suggest it could have reached over 12 meters (39 ft). Relative to the large and powerful hindlimbs, its three-fingered forelimbs were small, and the body was balanced by a long, heavy tail. It is classified as an allosaurid, a type of carnosaurian theropod dinosaur. The genus has a complicated taxonomy, and includes an uncertain number of valid species, the best known of which is A. fragilis. The bulk of Allosaurus remains have come from North America's Morrison Formation, with material also from Portugal and possibly Tanzania. It was known for over half of the 20th century as Antrodemus, but study of the copious remains from the Cleveland-Lloyd Dinosaur Quarry brought the name Allosaurus back to prominence, and established it as one of the best-known dinosaurs.

As the prominent large predator in the Morrison Formation, Allosaurus was at the top of the food chain, probably preying on contemporaneous large herbivorous dinosaurs. Potential prey included ornithopods, stegosaurids, and sauropods. While it is often thought of as preying on sauropod dinosaurs in groups, there is little evidence for cooperative social behavior in this genus, and individuals may have been aggressive toward each other instead. It may have attacked large prey by ambush, using its upper jaws like a hatchet. (see more...)

Massospondylus (meaning 'elongated vertebra') is a genus of prosauropod dinosaur from the early Jurassic Period (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by Sir Richard Owen in 1854 from remains found in South Africa, and is thus one of the first dinosaurs to have been named. Fossils have since been found in Lesotho, Zimbabwe, and other parts of South Africa. Further material from Arizona's Kayenta Formation, India, and Argentina has been assigned to this genus, but may not belong to Massospondylus.

The type, and only universally recognized species, is M. carinatus, although six other species have been named during the past 150 years. Prosauropod systematics have undergone numerous revisions during the last several years, and many scientists disagree where exactly Massospondylus lies on the dinosaur evolutionary tree. The family name Massospondylidae was once coined for the genus, but because knowledge of prosauropod relationships is in a state of flux, it is unclear which other dinosaurs—if any—belong in a natural grouping of massospondylids; several 2007 papers support the family's validity.

Although Massospondylus was long depicted as quadrupedal, a 2007 study found it to be bipedal. It was probably a herbivore, although it is speculated that the prosauropods may have been omnivorous. This animal, 4–6 meters (13–20 ft) long, had a long neck and tail, with a small head and slender body. On each of its forefeet, it bore a sharp thumb claw that was used in defense or feeding. Recent studies indicate Massospondylus grew steadily throughout its lifespan, possessed air sacs similar to those of birds, and may have cared for its young. (see more...)

Tarbosaurus (meaning 'terrifying lizard') is a genus of tyrannosaurid theropod dinosaur that flourished in Asia between 70 and 65 million years ago, near the end of the Late Cretaceous Period. Fossils have been recovered in Mongolia with more fragmentary remains found further afield in parts of China. Although many species have been named, modern paleontologists recognize only one, T bataar, as valid. Some experts contend that this species is actually an Asian representative of the North American genus Tyrannosaurus; if true, this would invalidate the genus Tarbosaurus altogether.

Tarbosaurus and Tyrannosaurus are considered closely related genera, even if they are not synonymous. Alioramus, also from Mongolia, is thought by some authorities to be the closest relative of Tarbosaurus. Like most known tyrannosaurids, Tarbosaurus was a large bipedal predator, weighing more than a ton and equipped with dozens of large, sharp teeth. It had a unique locking mechanism in its lower jaw and the smallest forelimbs relative to body size of all tyrannosaurids, renowned for their disproportionately tiny, two-fingered forelimbs.

Tarbosaurus lived in a humid floodplain criss-crossed by river channels. In this environment, it was an apex predator at the top of the food chain, probably preying on other large dinosaurs like the hadrosaur Saurolophus or the sauropod Nemegtosaurus. Tarbosaurus is very well-represented in the fossil record, known from dozens of specimens, including several complete skulls and skeletons. These remains have allowed scientific studies focusing on its phylogeny, skull mechanics, and brain structure. (see more...)

Gorgosaurus (meaning 'fierce lizard') is a genus of tyrannosaurid theropod dinosaur that lived in western North America during the Late Cretaceous Period, between about 77 and 74 million years ago. Fossil remains have been found in the Canadian province of Alberta and possibly the U.S. state of Montana. Paleontologists recognize only the type species, G. libratus, although other species have been erroneously referred to the genus.

Like most known tyrannosaurids, Gorgosaurus was a bipedal predator weighing more than a metric ton as an adult; dozens of large, sharp teeth lined its jaws, while its two-fingered forelimbs were comparatively small. Gorgosaurus was most closely related to Albertosaurus, and more distantly related to the larger Tyrannosaurus. Gorgosaurus and Albertosaurus are extremely similar, distinguished mainly by subtle differences in the teeth and skull bones. Some experts consider G. libratus to be a species of Albertosaurus; this would make Gorgosaurus a junior synonym of that genus.

Gorgosaurus lived in a lush floodplain environment along the edge of an inland sea. An apex predator, it was at the top of the food chain, preying upon abundant ceratopsids and hadrosaurs. In some areas, Gorgosaurus coexisted with another tyrannosaurid, Daspletosaurus. Though these animals were roughly the same size, there is some evidence of niche differentiation between the two. Gorgosaurus is the best-represented tyrannosaurid in the fossil record, known from dozens of specimens. These plentiful remains have allowed scientists to investigate its ontogeny, life history and other aspects of its biology. (see more...)

Amargasaurus was a sauropod dinosaur that lived in what is now Argentina from roughly 129 to 122 million years ago, during the Early Cretaceous epoch. The only known skeleton was discovered in 1984 and is virtually complete. Amargasaurus cazaui, the only species in the genus, was a large animal reaching 9 to 10 meters (30 to 33 feet) in length, with two parallel rows of tall spines down its neck and back. The spines, taller than in any other known sauropod, probably protruded as solitary structures supporting a keratinous sheath, and may have been used for display, combat, or defense. Alternatively, they might have formed a scaffold supporting a skin sail. A herbivore, Amargasaurus probably fed at mid-height. Discovered in sedimentary rocks of the La Amarga Formation, it is most closely related to the Late Jurassic genera Dicraeosaurus, Brachytrachelopan and Suuwassea. Together, these genera form the family Dicraeosauridae, with shorter necks and smaller body sizes than other sauropods. (Full article...)

Cope (left) and Marsh (right).

The Bone Wars is the name given to a period of intense fossil speculation and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope and Othniel Charles Marsh. The two paleontologists used underhanded methods to out-compete the other in the field, resorting to bribery, theft, and destruction of bones. The scientists also attacked each other in scientific publications, attempting to ruin the other's credibility and cut off his funding.

Originally colleagues who were civil to each other, Cope and Marsh became bitter enemies after several personal slights between them. Their pursuit of bones led them west to rich bone beds in Colorado, Nebraska, and Wyoming. From 1877 to 1892, both paleontologists used their wealth and influence to finance their own expeditions and to procure services and fossils from dinosaur hunters. By the end of the Bone Wars, both men exhausted their funds in fueling their intense rivalry.

Cope and Marsh were financially and socially ruined by their efforts to disgrace each other, but their contributions to science and the field of paleontology were massive; the scientists left behind tons of unopened boxes of fossils on their deaths. The feud between the two men led to over 142 new species of dinosaurs being discovered and described. The products of the Bone Wars resulted in an increase in knowledge of ancient life, and sparked the public's interest in dinosaurs, leading to continued fossil excavation in North America in the decades to come. Several historical books and fictional adaptations have also been published about this period of intense paleontological activity. (see more...)

Edmontosaurus (meaning 'Edmonton [Formation] lizard') is a genus of crestless duck-billed dinosaur. The fossils of this animal have been found in rocks of western North America that date from the late Campanian stage to the end of the Maastrichtian stage of the Cretaceous Period, between 73 and 65.5 million years ago. It was one of the last non-avian dinosaurs, and lived alongside Triceratops and Tyrannosaurus shortly before the Cretaceous–Paleogene extinction event. Edmontosaurus was one of the largest hadrosaurids, measuring up to 13 meters (43 ft) long and weighing around 4.0 metric tons (4.4 short tons). It is known from several well-preserved specimens that include not only bones, but in some cases extensive skin impressions and possible gut contents.

Edmontosaurus has a lengthy and complicated taxonomic history dating to the late 19th century. Various species classified with genera such as Claosaurus, Thespesius, Trachodon, and the well-known but now defunct genus Anatosaurus are now regarded as belonging to Edmontosaurus. The first fossils named Edmontosaurus were discovered in southern Alberta, Canada, in what used to be called the lower Edmonton Formation. The type species, E. regalis, was named by Lawrence Lambe in 1917, although several other species that are now classified in Edmontosaurus were named earlier. The best known of these is E. annectens, originally named by Othniel Charles Marsh in 1892 as Claosaurus annectens and known for many years as Anatosaurus annectens.

Edmontosaurus was widely distributed across western North America. The distribution of Edmontosaurus fossils suggests that it preferred coasts and coastal plains. It was an herbivore that could move on both two legs and four. Because it is known from several bone beds, Edmontosaurus is thought to have lived in groups, and may have been migratory as well. The wealth of fossils has allowed researchers to study its paleobiology in detail, including its brain, how it may have fed, and its injuries and pathologies, such as evidence for a tyrannosaur attack on one edmontosaur specimen. (see more...)

Herrerasaurus (meaning 'Herrera's lizard', after the name of the rancher who discovered the first fossil of the animal) was one of the earliest dinosaurs. All known specimens of this carnivore have been discovered in rocks of late Ladinian age (middle Triassic according to the ICS, dated to 231.4 million years ago) in northwestern Argentina. The type species, Herrerasaurus ischigualastensis, was described by Osvaldo Reig in 1963 and is the only species assigned to the genus. The names Ischisaurus and Frenguellisaurus are synonymous with Herrerasaurus. For many years, the classification of Herrerasaurus was unclear, as the animal was initially known from very fragmentary remains; it has been hypothesized to be a basal theropod, a basal sauropodomorph, a basal saurischian, or not a dinosaur at all. However, with the discovery of a mostly complete skeleton and skull in 1988, Herrerasaurus has been classified as either an early theropod or an early saurischian in at least five recent reviews of theropod evolution. It was a member of the Herrerasauridae, a group of similar animals which were among the earliest of the dinosaurian evolutionary radiation. (see more...)

Edward Drinker Cope (born July 28, 1840 – died April 12, 1897) was an American paleontologist and comparative anatomist, as well as a noted herpetologist and ichthyologist. Born to a wealthy Quaker family, Cope distinguished himself as a child prodigy interested in science; he published his first scientific paper at the age of nineteen. Though his father tried to raise Cope as a gentleman farmer, he eventually acquiesced to his son's scientific aspirations. Cope married his cousin and had one child; the family moved from Philadelphia to Haddonfield, New Jersey, although Cope would maintain a residence and museum in Philadelphia in his later years.

Cope had little formal scientific training, and he eschewed a teaching position for field work. He made regular trips to the American West prospecting in the 1870s and 1880s, often as a member of United States Geological Survey teams. A personal feud between Cope and paleontologist Othniel Charles Marsh led to a period of intense fossil-finding competition now known as the Bone Wars. Cope's financial fortunes soured after failed mining ventures in the 1880s, forcing him to sell off much of his fossil collection. He experienced a resurgence in his career toward the end of his life before dying of unidentified causes on April 12, 1897.

Cope's scientific pursuits nearly bankrupted him, but his contributions helped to define the field of American paleontology. He was a prodigious writer, with 1,400 papers published over his lifetime, although his rivals would debate the accuracy of his rapidly published works. He discovered, described, and named more than 1,000 vertebrate species including hundreds of fishes and dozens of dinosaurs. His proposals on the origin of mammalian molars and for the gradual enlargement of mammalian species over geologic time ("Cope's Law") are notable among his theoretical contributions. (see more...)

Dinosaurs are vertebrates that have ranged from reptile-like to bird-like. Dinosaurs dominated the terrestrial ecosystem for over 160 million years from around 230 million years ago until 65 million years ago, at the end of the Cretaceous period, when all non-avian dinosaurs became extinct. Knowledge about dinosaurs comes from both fossil and non-fossil records, including fossilized bones, feces, trackways, gastroliths, feathers, impressions of skin, internal organs and soft tissues. Dinosaur remains have been found on every continent on Earth, including Antarctica, showing that all land masses were at one time connected in a supercontinent called Pangaea. The ongoing dinosaur renaissance began in the 1970s and was triggered, in part, by John Ostrom's discovery of Deinonychus, an active, vicious predator that may have been warm-blooded (homeothermic), in contrast to the prevailing image of dinosaurs as sluggish and cold-blooded. Vertebrate paleontology has also become a global science, with major new discoveries in previously unexploited regions, most significantly the amazingly well-preserved feathered dinosaurs in China, which have further solidified the link between dinosaurs and their living descendants, modern birds. (see more...)