Eels are ray-finned fish belonging to the order Anguilliformes (//), which consists of eight suborders, 19 families, 111 genera, and about 800 species. Eels undergo considerable development from the early larval stage to the eventual adult stage, and most are predators.
L. S. Berg, 1943
The term "eel" is also used for some other eel-shaped fish, such as electric eels (genus Electrophorus), spiny eels (family Mastacembelidae), swamp eels (family Synbranchidae), and deep-sea spiny eels (family Notacanthidae). These other clades, however, evolved their eel-like shapes independently from the true eels. Eels live both in salt water and fresh water and some species are catadromous.
Eels are elongated fish, ranging in length from 5 cm (2 in) in the one-jawed eel (Monognathus ahlstromi) to 4 m (13 ft) in the slender giant moray. Adults range in weight from 30 g (1 oz) to well over 25 kg (55 lb). They possess no pelvic fins, and many species also lack pectoral fins. The dorsal and anal fins are fused with the caudal fin, forming a single ribbon running along much of the length of the animal. Eels swim by generating waves which travel the length of their bodies. They can swim backwards by reversing the direction of the wave.
Most eels live in the shallow waters of the ocean and burrow into sand, mud, or amongst rocks. A majority of eel species are nocturnal, thus are rarely seen. Sometimes, they are seen living together in holes, or "eel pits". Some species of eels also live in deeper water on the continental shelves and over the slopes deep as 4,000 m (13,000 ft). Only members of the Anguilla regularly inhabit fresh water, but they, too, return to the sea to breed.
The heaviest true eel is the European conger. The maximum size of this species has been reported as reaching a length of 3 m (10 ft) and a weight of 110 kg (240 lb). Other eels are longer, but do not weigh as much, such as the slender giant moray which reaches 4 m (13 ft).
Eels begin life as flat and transparent larvae, called leptocephali. Eel larvae drift in the surface waters of the sea, feeding on marine snow, small particles that float in the water. Eel larvae then metamorphose into glass eels and then become elvers before finally seeking out their juvenile and adult habitats. Many eels remain in the sea throughout their lives, but freshwater elvers of eels in the family Anguillidae travel upstream and are forced to climb up obstructions, such as weirs, dam walls, and natural waterfalls.
Eel eggs hatch firstly into the leptocephalus larval stage
Larval eels become glass eels as they transition from the ocean to fresh water
A number of classifications of eels exist; some, such as FishBase, divide eels into 20 families, whereas other classifications such as ITIS and Systema Naturae 2000 include additional eel families, which are noted below the family with which they are synonymized in the FishBase system.
Suborders and familiesEdit
Taxonomy based on Nelson, Grande and Wilson 2016.
- Suborder Protanguilloidei
- Family Protanguillidae
- Suborder Synaphobranchoidei
- Family Synaphobranchidae (cutthroat eels) [incl. Dysommidae, Nettodaridae, and Simenchelyidae]
- Suborder Muraenoidei
- Suborder Chlopsoidei
- Family Chlopsidae (false morays)
- Suborder Congroidei
- Suborder Moringuoidei
- Family Moringuidae (spaghetti eels)
- Suborder Saccopharyngoidei
- Suborder Anguilloidei
Phylogeny based on Johnson et al. 2012.
|Main commercial species|
|Common name||Scientific name||Maximum
|American eel||Anguilla rostrata (Lesueur, 1817)||152 cm||50 cm||7.33 kg||43 years||3.7|||||| |
|European eel||Anguilla anguilla (Linnaeus, 1758)||150 cm||35 cm||6.6 kg||88 years||3.5|||||||| |
|Japanese eel||Anguilla japonica Temminck & Schlegel, 1846||150 cm||40 cm||1.89 kg||3.6|||||||| |
|Short-finned eel||Anguilla australis Richardson, 1841||130 cm||45 cm||7.48 kg||32 years||4.1|||||| |
Use by humansEdit
Freshwater eels (unagi) and marine eels (conger eel, anago) are commonly used in Japanese cuisine; foods such as unadon and unajū are popular, but expensive. Eels are also very popular in Chinese cuisine, and are prepared in many different ways. Hong Kong eel prices have often reached 1000 HKD (128.86 US Dollars) per kg, and once exceeded 5000 HKD per kg. In India, eels are popularly eaten in the Northeast. Freshwater eels, known as Kusia in Assamese, are eaten with curry, often with herbs. The European eel and other freshwater eels are mostly eaten in Europe, the United States, and other places. Today, the European eel is considered critically endangered. A traditional east London food is jellied eels, although the demand has significantly declined since World War II. The Spanish cuisine delicacy angulas consists of elver (young eels) sautéed in olive oil with garlic; elvers usually reach prices of up to 1000 euro per kg. New Zealand longfin eel is a traditional Māori food in New Zealand. In Italian cuisine, eels from the Valli di Comacchio, a swampy zone along the Adriatic coast, are especially prized, along with freshwater eels of Bolsena Lake and pond eels from Cabras, Sardinia. In northern Germany, the Netherlands, the Czech Republic, Poland, Denmark, and Sweden, smoked eel is considered a delicacy.
Eel blood is toxic to humans and other mammals, but both cooking and the digestive process destroy the toxic protein. The toxin derived from eel blood serum was used by Charles Robert Richet in his Nobel Prize-winning research which discovered anaphylaxis (by injecting it into dogs and observing the effect). The poison used by Richet was actually obtained from sea anemones.
Eelskin leather is highly prized. It is very smooth and exceptionally strong. It does not come from eels. It comes from the Pacific hagfish, a jawless fish which is also known as the slime eel.
The large lake of Almere, which existed in the early Medieval Netherlands, got its name from the eels which lived in its water (the Dutch word for eel is aal or ael, so: "ael mere" = "eel lake"). The name is preserved in the new city of Almere in Flevoland, given in 1984 in memory of this body of water on whose site the town is located.
Eel fishing in Nazi-era Danzig plays an important role in Günter Grass' novel The Tin Drum. The cruelty of humans to eels is used as a metaphor for Nazi atrocities, and the sight of eels being killed by a fisherman triggers the madness of the protagonist's mother.
Sinister implications of eels fishing are also referenced in Jo Nesbø's Cockroaches, the second book of the Harry Hole detective series. The book's background includes a Norwegian village where eels in the nearby sea are rumored to feed on the corpses of drowned humans, making the eating of these eels verge on cannibalism.
The English name "eel" descends from Old English ǣl, Common Germanic *ēlaz. Also from the common Germanic are West Frisian iel, Dutch aal, German Aal, and Icelandic áll. Katz (1998) identifies a number of Indo-European cognates, among them the second part of the Latin word for eels, anguilla, attested in its simplex form illa (in a glossary only), and the Greek word for "eel", egkhelys (the second part of which is attested in Hesychius as elyes). The first compound member, anguis ("snake"), is cognate to other Indo-European words for "snake" (compare Old Irish escung "eel", Old High German unc "snake", Lithuanian angìs, Greek ophis, okhis, Vedic Sanskrit áhi, Avestan aži, Armenian auj, iž, Old Church Slavonic *ǫžь, all from Proto-Indo-European *h₁ogʷʰis).The word also appears in the Old English word for "hedgehog," which is igil (meaning "snake eater"), and perhaps in the egi- of Old High German egidehsa "wall lizard".
According to this theory, the name Bellerophon (Βελλεροφόντης, attested in a variant Ἐλλεροφόντης in Eustathius of Thessalonica), is also related, translating to "the slayer of the serpent" (ahihán). On this theory, the ελλερο- is an adjective form of an older word, ελλυ, meaning "snake", which is directly comparable to Hittite ellu-essar- "snake pit". This myth likely came to Greece via Anatolia. In the Hittite version of the myth, the dragon is called Illuyanka: the illuy- part is cognate to the word illa, and the -anka part is cognate to angu, a word for "snake". Since the words for "snake" (and similarly shaped animals) are often subject to taboo in many Indo-European (and non-Indo-European) languages, no unambiguous Proto-Indo-European form of the word for eel can be reconstructed. It may have been *ēl(l)-u-, *ēl(l)-o-, or something similar.
Timeline of generaEdit
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