Homo erectus (meaning 'upright man') were a species of archaic humans from the Pleistocene, earliest occurrence about 2 mya. They are proposed to be the direct ancestors to several human species, such as H. heidelbergensis, H. antecessor, Neanderthals, Denisovans, and modern humans. As a chronospecies, the time of its disappearance is thus a matter of contention or even convention. There are also several proposed subspecies with varying levels of recognition.
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H. erectus appear to have been much more similar to modern humans than to their ancestors, the australopithecines, with a more humanlike gait, body proportions, height, and brain capacity. H. erectus are associated with the Acheulean stone tool industry, and is thought to be the earliest human ancestor capable of starting fires, speech, hunting and gathering in coordinated groups, caring for injured or sick group members, and possibly art-making.
The first remains, Java Man, were described by Dutch anatomist Eugène Dubois in 1893, who set out to look for the "missing link" between apes and humans in Southeast Asia, because he believed gibbons to be the closest living relatives to humans in accordance with the "Out of Asia" hypothesis. H. erectus was the first fossil hominin found as a result of a directed expedition.
Excavated from the bank of the Solo River at Trinil, East Java, he first allocated the material to a genus of fossil chimpanzees as Anthropopithecus erectus, then the following year assigned it to a new genus as Pithecanthropus erectus (the genus name had been coined by Ernst Haeckel in 1868 for the hypothetical link between humans and fossil Apes). The species name erectus was given because the femur suggested that Java Man had been bipedal and walked upright. However, few scientists recognized it as a "missing link", and, consequently, Dubois' discovery had been largely disregarded.
In 1921, two teeth from Zhoukoudian, China discovered by Johan Gunnar Andersson had prompted widely publicized interest. When describing the teeth, Davidson Black named it a new species Sinanthropus pekinensis from Ancient Greek Σίνα sino- "China" and Latin pekinensis "of Peking". Subsequent excavations uncovered about 200 human fossils from more than 40 individuals including five nearly complete skullcaps. Franz Weidenreich provided much of the detailed description of this material in several monographs published in the journal Palaeontologica Sinica (Series D). Nearly all of the original specimens were lost during World War II during an attempt to smuggle them out of China for safekeeping. However, casts were made by Weidenreich, which exist at the American Museum of Natural History in New York City and at the Institute of Vertebrate Paleontology and Paleoanthropology in Beijing.
Similarities between Java Man and Peking Man led Ernst Mayr to rename both as Homo erectus in 1950. Throughout much of the 20th century, anthropologists debated the role of H. erectus in human evolution. Early in the century, due in part to the discoveries at Java and Zhoukoudian, the belief that modern humans first evolved in Asia was widely accepted. A few naturalists—Charles Darwin the most prominent among them—theorized that humans' earliest ancestors were African. Darwin pointed out that chimpanzees and gorillas, humans' closest relatives, evolved and exist only in Africa.
It has been proposed that H. erectus evolved from H. habilis about 2 mya, though this has been called into question because they coexisted for at least a half a million years. Alternatively, a group of H. habilis may have been reproductively isolated, and only this group developed into H. erectus (cladogenesis).
Because the earliest remains of H. erectus are found in both Africa and East Asia (in China as early as 2.1 mya), it is debated where H. erectus evolved. A 2011 study suggested that it was H. habilis who reached West Asia, that early H. erectus developed there, and that early H. erectus would then have dispersed from West Asia to East Asia (Peking Man), Southeast Asia (Java Man), back to Africa (Homo ergaster), and to Europe (Tautavel Man), eventually evolving into modern humans in Africa. Others have suggested that H. erectus/H. ergaster developed in Africa, where it eventually evolved into modern humans.
- Homo erectus bilzingslebenensis (0.37 Ma)
- Homo erectus erectus (Java Man, 1.6–0.5 Ma)
- Homo erectus ergaster (1.9–1.4 Ma)
- Homo erectus georgicus (1.8–1.6 Ma)
- Homo erectus heidelbergensis (0.7–0.3 Ma), evidence exists of archaic human presence in Europe before 1 mya, as early as 1.6 mya.
- Homo erectus lantianensis (Lantian Man, 1.6 Ma)
- Homo erectus nankinensis (Nanjing Man, 0.6 Ma)
- Homo erectus palaeojavanicus (Meganthropus, 1.4–0.9 Ma)
- Homo erectus pekinensis (Peking Man, 0.7 Ma)
- Homo erectus soloensis (Solo Man, 0.546–0.143 Ma)
- Homo erectus tautavelensis (Tautavel Man, 0.45 Ma)
- Homo erectus yuanmouensis (Yuanmou Man)
Since its discovery in 1893 (Java man), there has been a trend in palaeoanthropology of reducing the number of proposed species of Homo, to the point where H. erectus includes all early (Lower Paleolithic) forms of Homo sufficiently derived from H. habilis and distinct from early H. heidelbergensis (in Africa also known as H. rhodesiensis). It is sometimes considered as a wide-ranging, polymorphous species.
Due to such a wide range of variation, it has been suggested that the ancient H. rudolfensis and H. habilis should be considered early varieties of H. erectus. The primitive H. e. georgicus from Dmanisi, Georgia has the smallest brain capacity of any known Pleistocene hominin (about 600 cc), and its inclusion in the species would greatly expand the range of variation of H. erectus to perhaps include species as H. rudolfensis, H. gautengensis, H. ergaster, and perhaps H. habilis. However, a 2015 study suggested that H. georgicus represents an earlier, more primitive species of Homo derived from an older dispersal of hominins from Africa, with H. ergaster/erectus possibly deriving from a later dispersal. H. georgicus is sometimes not even regarded as H. erectus.
It is debated whether the African H. e. ergaster is a separate species (and that H. erectus evolved in Asia, then migrated to Africa), or is the African form (sensu lato) of H. erectus (sensu stricto). In the latter, H. ergaster has also been suggested to represent the immediate ancestor of H. erectus. It has also been suggested that H. ergaster instead of H. erectus, or some hybrid between the two, was the immediate ancestor of other archaic humans and modern humans.
Descendents and synonymsEdit
Homo erectus is the most long-lived species of Homo, having survived for over a million years. By contrast, Homo sapiens emerged about a quarter million years ago.
- African H. erectus candidates
- Eurasian H. erectus candidates:
- Homo floresiensis
- Homo sapiens candidates
- Homo neanderthalensis (or H. s. neanderthalensis)
- Homo denisova (or H. s. denisova or Homo sp. Altai, and Homo sapiens subsp. Denisova)
- Homo rhodesiensis (or H. s. rhodensis)
- Homo heidelbergensis (or H. s. heidelbergensis)
- Homo sapiens idaltu
- the Narmada fossil, discovered in 1982 in Madhya Pradesh, India, was at first suggested as H. erectus (Homo erectus narmadensis) but later recognized as H. sapiens.
If considering Homo erectus in its strict sense (that is, as referring to only the Asian variety) no consensus has been reached as to whether it is ancestral to H. sapiens or any later human species. Similarly, H. antecessor and sisters, including modern humans, appear to have emerged specifically as sister of, for example, the Asian variety of H. erectus. Moreover, some late H. erectus varieties may have introgressed into the Denisovans, which then later introgressed into H. sapiens. However, it is conventional to label European archaic humans as H. heidelbergensis, the immediate predecessor of Neanderthals.
Meganthropus refers to a group of fossils found in Java, dated to between 1.4 and 0.9 Mya, which are tentatively grouped with H. erectus, at least in the wider sense of the term in which "all earlier Homo populations that are sufficiently derived from African early Homo belong to H. erectus", although older literature has placed the fossils outside of Homo altogether.
One of the features distinguishing H. erectus from H. sapiens is the size difference in teeth. H. erectus has large teeth while H. sapiens have smaller teeth. One theory for the reason of H. erectus having larger teeth is because of the requirements for eating raw meat instead of cooked meat like H. sapiens.
Foot tracks found in 2009 in Kenya confirmed that the gait of H. erectus was heel-to-toe, more like modern humans than australopithecines.
H. erectus fossils show a cranial capacity greater than that of Homo habilis (although the Dmanisi specimens have distinctively small crania): the earliest fossils show a cranial capacity of 850 cm³, while later Javan specimens measure up to 1100 cm³, overlapping that of H. sapiens.
The frontal bone is less sloped and the dental arcade smaller than that of the australopithecines; the face is [clarify] than australopithecines or H. habilis, with large brow-ridges and less prominent cheekbones.
H. erectus is estimated to have had an average height of 1.8 m (5.9 ft), similar to modern humans. Males were about 25% larger than females, a difference slightly greater than that seen in modern humans, but less than that of Australopithecus. H. erectus is thought to have been quite slender, with long arms and legs.
Throwing performance may have been an important mode for early hunting and defense in the genus Homo, linked to several anatomical shifts. H. erectus may have had a shoulder configuration similar to humans, laterally-facing with respect to the torso. This suggests that the ability for high speed throwing can be dated back to nearly two million years ago.
H. erectus is credited with inventing the Acheulean stone tool industry, succeeding the Oldowan industry, and were the first to make hand axes out of stone. Though larger and heavier, these hand axes had sharper, chiseled edges, likely used in butchering. Such tools could indicate H. erectus had the ability of foresight and could plan steps ahead in order to create them.
The earliest record of Achuelean technology comes from West Turkana, Kenya 1.76 mya. Oldowan lithics are also known from the site, and the two seemed to coexist for some time. The earliest records of Achuelean technology outside of Africa date to no older than 1 mya, indicating it only became widespread after some secondary H. erectus dispersal from Africa.
It has been suggested that the Asian H. erectus may have been the first humans to use rafts to travel over bodies of water, including oceans. The oldest stone tool found in Turkey reveals that hominins passed through the Anatolian gateway from western Asia to Europe approximately 1.2 million years ago—much earlier than previously thought.
It is unknown if they wore clothes and had tools such as bowls and utensils.
Use of fireEdit
A site at Bnot Ya'akov Bridge, Israel is reported to show evidence that H. erectus or H. ergaster controlled fire there between 790,000 and 690,000 years ago; to date this claim has been widely accepted. Some evidence is found that H. erectus was controlling fire less than 250,000 years ago. Evidence also exists that H. erectus were cooking their food as early as 500,000 years ago. Re-analysis of burnt bone fragments and plant ashes from the Wonderwerk Cave, South Africa, could be considered evidence of cooking as early as 1 mya.
Several East African sites show some evidence of early fire usage. At Chesowanja, there is evidence of 1.42 Ma fire-hardened clay heated to about 400 °C (752 °F), consistent with campfires. At two sites in Koobi Fora, there is a reddening of sediment associated with heating the material to 200–400 °C (392–752 °F). There is a hearth-like depression with traces of charcoal at Olorgesailie, but this could have resulted from a natural bush fire.
In Gadeb, Ethiopia, fragments of welded tuff that appeared to have been burned or scorched were found alongside Acheulean artifacts. However, such re-firing of the rocks may have been caused by local volcanic activity. In the Middle Awash River Valley, cone-shaped depressions of reddish clay were found that could have been created only by temperatures of 200 °C (392 °F) or greater. These depressions may have been burnt tree stumps, and the fire was made far from the habitation site. Burnt stones are found in the Awash Valley, but naturally burnt (volcanic) welded tuff is also found in the area.
Remains of H. erectus have more consistently been found in lake beds and in stream sediments. This suggests that H. erectus also lived in open encampments near streams and lakes.
An engraved shell with geometric markings could possibly be evidence of the earliest art-making, dating back to 546–436 kya. Art-making capabilities could be considered evidence of symbolic thinking, which is associated with modern cognition and behavior.
Homo erectus was probably the first hominin to live in a hunter-gatherer society, and anthropologists such as Richard Leakey believe that H. erectus was socially more like modern humans than australopithecines. Likewise, increased cranial capacity generally correlates to the more sophisticated tools H. erectus are associated with.
The discovery of Turkana boy (H. ergaster) in 1984 evidenced that, despite its Homo sapiens-like anatomy, ergaster may not have been capable of producing sounds comparable to modern human speech. It likely communicated in a proto-language, lacking the fully developed structure of modern human language, but more developed than the non-verbal communication used by chimpanzees. This inference is challenged by the find in Dmanisi, Georgia, of a vertebra (at least 150,000 years earlier than the Turkana Boy) that reflects vocal capabilities within the range of H. sapiens. Both brain size and the presence of the Broca's area also support the use of articulate language.
H. erectus were probably the first human ancestor to live in small, familiar band-societies similar to modern hunter-gatherers, and are thought to be the first to hunt in coordinated groups, to use complex tools, and to care for infirm or weak companions.
The lower cave of the Zhoukoudian cave, China, is one of the most important archaeological sites worldwide. There have been remains of 45 homo erectus individuals found and thousands of tools recovered. Most of these remains were lost during World War 2, with the exception of two postcranial elements that were rediscovered in China in 1951 and four human teeth from 'Dragon Bone Hill'.
New evidence has shown that Homo erectus does not have uniquely thick vault bones, like what was previously thought. Testing showed that neither Asian or African Homo erectus had uniquely large vault bones.
Some of the major Homo erectus fossils:
- Indonesia (island of Java): Trinil 2 (holotype), Sangiran collection, Sambungmachan collection, Ngandong collection
- China ("Peking Man"): Lantian (Gongwangling and Chenjiawo), Yunxian, Zhoukoudian, Nanjing, Hexian
- Kenya: KNM ER 3883, KNM ER 3733
- Indriati, E; Swisher, CC 3rd; Lepre, C; Quinn, RL; Suriyanto, RA; Hascaryo, AT; Grün, R; Feibel, CS; Pobiner, BL; Aubert, M; Lees, W; Antón, SC (2011). "The age of the 20 meter Solo River terrace, Java, Indonesia and the survival of Homo erectus in Asia". PLOS ONE. 6: e21562. doi:10.1371/JOURNAL.PONE.0021562. PMC 3126814. PMID 21738710.
- based on numerous fossil remains of H. erectus. Museum of Prehistory Tautavel, France (2008 photograph)
- Klein, R. (1999). The Human Career: Human Biological and Cultural Origins. Chicago: University of Chicago Press, ISBN 0226439631.
- Wood, Bernard (2011). "Did early Homo migrate "out of" or "in to" Africa?". Proceedings of the National Academy of Sciences. 108: 10375–10376. doi:10.1073/pnas.1107724108.
- Swisher, Curtis & Lewin 2000, p. 70.
- Reconstruction by John Gurche (2010), Smithsonian Museum of Natural History, based on KNM ER 3733 and 992. Abigail Tucker, "A Closer Look at Evolutionary Faces", Smithsonian.com, 25 February 2010.
- "The First Knock at the Door". Peking Man Site Museum.
In the summer of 1921, Dr. J.G. Andersson and his companions discovered this richly fossiliferous deposit through the local quarry men's guide. During examination he was surprised to notice some fragments of white quartz in tabus, a mineral normally foreign in that locality. The significance of this occurrence immediately suggested itself to him and turning to his companions, he exclaimed dramatically "Here is primitive man, now all we have to do is find him!"
- "Review of the History". Peking Man Site Museum.
During 1927–1937, abundant human and animal fossils as well as artefact were found at Peking Man Site, it made the site to be the most productive one of the Homo erectus sites of the same age all over the world. Other localities in the vicinity were also excavated almost at the same time.
- Darwin, Charles R. (1871). The Descent of Man and Selection in Relation to Sex. John Murray. ISBN 978-0-8014-2085-6.
- Reconstruction by W. Schnaubelt & N. Kieser (Atelier Wild Life Art), 2006, Westfälisches Museum für Archäologie, Herne, Germany.
- F. Spoor; M.G. Leakey; P.N. Gathogo; F.H. Brown; S.C. Antón; I. McDougall; C. Kiarie; F.K. Manthi; L.N. Leakey (9 August 2007). "Implications of new early Homo fossils from Ileret, east of Lake Turkana, Kenya". Nature. 448 (7154): 688–691. Bibcode:2007Natur.448..688S. doi:10.1038/nature05986. PMID 17687323.
A partial maxilla assigned to H. habilis reliably demonstrates that this species survived until later than previously recognized, making an anagenetic relationship with H. erectus unlikely
- Zhu Zhaoyu (朱照宇); Dennell, Robin; Huang Weiwen (黄慰文); Wu Yi (吴翼); Qiu Shifan (邱世藩); Yang Shixia (杨石霞); Rao Zhiguo (饶志国); Hou Yamei (侯亚梅); Xie Jiubing (谢久兵); Han Jiangwei (韩江伟); Ouyang Tingping (欧阳婷萍) (2018). "Hominin occupation of the Chinese Loess Plateau since about 2.1 million years ago". Nature. 559 (7715): 608–612. Bibcode:2018Natur.559..608Z. doi:10.1038/s41586-018-0299-4. ISSN 0028-0836. PMID 29995848.
- Barras, Colin (2018). "Tools from China are oldest hint of human lineage outside Africa". Nature. doi:10.1038/d41586-018-05696-8. ISSN 0028-0836.
- Denell, R. (11 July 2018). "Hominin occupation of the Chinese Loess Plateau since about 2.1 million years ago". US National Library of Medicine National Institutes of Health. 559 (7715): 608–612. doi:10.1038/s41586-018-0299-4. PMID 29995848.
- Hao, L; Chao Rong, L; Kuman, K (2017). "Longgudong, an Early Pleistocene site in Jianshi, South China, with stratigraphic association of human teeth and lithics". Science China Earth. 60 (3): 452–462.
- Ferring, R.; Oms, O.; Agusti, J.; Berna, F.; Nioradze, M.; Shelia, T.; Tappen, M.; Vekua, A.; Zhvania, D.; Lordkipanidze, D. (2011). "Earliest human occupations at Dmanisi (Georgian Caucasus) dated to 1.85-1.78 Ma". Proceedings of the National Academy of Sciences. 108 (26): 10432–10436. Bibcode:2011PNAS..10810432F. doi:10.1073/pnas.1106638108. PMC 3127884. PMID 21646521.
- Augusti, Jordi; Lordkipanidze, David (June 2011). "How "African" was the early human dispersal out of Africa?". Quaternary Science Reviews. 30 (11–12): 1338–1342. Bibcode:2011QSRv...30.1338A. doi:10.1016/j.quascirev.2010.04.012.
- G. Philip Rightmire (1998). "Human Evolution in the Middle Pleistocene: The Role of Homo heidelbergensis". Evolutionary Anthropology. 6 (6): 218–227. doi:10.1002/(sici)1520-6505(1998)6:6<218::aid-evan4>3.0.co;2-6.
- Asfaw B, Gilbert WH, Beyene Y, Hart WK, Renne PR, WoldeGabriel G, Vrba ES, White TD (June 2002). "Remains of Homo erectus from Bouri, Middle Awash, Ethiopia". Nature. 416 (6878): 317–320. doi:10.1038/416317a. PMID 11907576.
- Moore, Matthew (8 July 2010). "Norfolk earliest known settlement in northern Europe". The Daily Telegraph. London. Retrieved 8 July 2010.
- Ghosh, Pallab (7 July 2010). "Humans' early arrival in Britain". BBC. Retrieved 8 July 2010.
- Jones, Tim (17 December 2009). "Lithic Assemblage Dated to 1.57 Million Years Found at Lézignan-la-Cébe, Southern France «". Anthropology.net. Retrieved 21 June 2012.
- Ciochon, RL (2009). "The mystery ape of Pleistocene Asia". Nature. 459: 910–911. doi:10.1038/459910a. PMID 19536242.
- Kaifu, Y.; et al. (2005). "Taxonomic affinities and evolutionary history of the Early Pleistocene hominids of Java: dentognathic evidence". Am. J. Phys. Anthropol. 128: 709–726. doi:10.1002/ajpa.10425.
- Skull suggests three early human species were one : Nature News & Comment
- David Lordkipanidze, Marcia S. Ponce de Leòn, Ann Margvelashvili, Yoel Rak, G. Philip Rightmire, Abesalom Vekua, Christoph P.E. Zollikofer (18 October 2013). "A Complete Skull from Dmanisi, Georgia, and the Evolutionary Biology of Early Homo". Science. 342 (6156): 326–331. Bibcode:2013Sci...342..326L. doi:10.1126/science.1238484. PMID 24136960.CS1 maint: multiple names: authors list (link)
- Switek, Brian (17 October 2013). "Beautiful Skull Spurs Debate on Human History". National Geographic. Retrieved 22 September 2014.
- Ian Sample (17 October 2013). "Skull of Homo erectus throws story of human evolution into disarray". The Guardian.
- Giumares SW, Merino CL (September 2015). "Dmanisi hominin fossils and the problem of multiple species in the early Homo genus" (PDF). Nexus: The Canadian Student Journal of Anthropology. 23.
- Argue, Debbie; Groves, Colin P.; Lee, Michael S.Y.; Jungers, William L. (23 February 2017). "The affinities of Homo floresiensis based on phylogenetic analyses of cranial, dental, and postcranial characters". Journal of Human Evolution. 107: 107–133. doi:10.1016/j.jhevol.2017.02.006. PMID 28438318.
- Lordkipanidze, David (4 October 2018), "Dmanisi", in Trevathan, Wenda; Cartmill, Matt; Dufour, Dana; Larsen, Clark (eds.), The International Encyclopedia of Biological Anthropology, 3 Volume Set, The International Encyclopedia of Biological Anthropology, John Wiley & Sons, Inc., pp. 1–4, doi:10.1002/9781118584538.ieba0139, ISBN 9781118584422
- Baab K (December 2015). "Defining Homo erectus". Handbook of Paleoanthropology (2 ed.): 2189–2219. doi:10.1007/978-3-642-39979-4_65 (inactive 4 December 2019).
- Tattersall, Ian and Jeffrey Schwartz (2001). Extinct Humans. Boulder, Colorado: Westview/Perseus. ISBN 978-0-8133-3482-0.[page needed]
- There was long-standing uncertainty whether H. floresiensis should be considered close to H. erectus, close to H. sapiens, or an altogether separate species. In 2017, it was suggested on morphological grounds that H. floresiensis is a sister species to either H. habilis or to a minimally habilis-erectus-ergaster-sapiens clade, and its line much more ancient than Homo erectus itself. Argue, Debbie; Groves, Colin P. (21 April 2017). "The affinities of Homo floresiensis based on phylogenetic analyses of cranial, dental, and postcranial characters". Journal of Human Evolution. 107: 107–133. doi:10.1016/j.jhevol.2017.02.006. PMID 28438318.
- Kennedy, Kenneth A.R.; Sonakia, Arun; Chiment, John; Verma, K.K. (1991). "Is the Narmada hominid an Indian Homo erectus?". American Journal of Physical Anthropology. 86 (4): 475–496. doi:10.1002/ajpa.1330860404. PMID 1776655.
- Dembo, Mana; Radovčić, Davorka; Garvin, Heather M.; Laird, Myra F.; Schroeder, Lauren; Scott, Jill E.; Brophy, Juliet; Ackermann, Rebecca R.; Musiba, Chares M.; de Ruiter, Darryl J.; Mooers, Arne Ø. (29 April 2016). "The evolutionary relationships and age of Homo naledi: An assessment using dated Bayesian phylogenetic methods". Journal of Human Evolution. 97: 17–26. doi:10.1016/j.jhevol.2016.04.008. PMID 27457542.
- Lao, Oscar; Bertranpetit, Jaume; Mondal, Mayukh (16 January 2019). "Approximate Bayesian computation with deep learning supports a third archaic introgression in Asia and Oceania". Nature Communications. 10 (1): 246. Bibcode:2019NatCo..10..246M. doi:10.1038/s41467-018-08089-7. ISSN 2041-1723. PMC 6335398. PMID 30651539.
- Krantz, G.S. (1975). "An explanation for the diastema of Javan erectus Skull IV". In: Paleoanthropology, Morphology and Paleoecology. La Hague: Mouton, 361–372.
- Homo erectus. (2018). Britannica Online Academic Edition, Encyclopædia Britannica, Inc.
- "A footprint in the sands of time". The Economist. 26 February 2009. Retrieved 22 December 2017.
- Swisher, Carl Celso III; Curtis, Garniss H. and Lewin, Roger (2002) Java Man, Abacus, ISBN 0-349-11473-0.
- Bryson, Bill (2005). A Short History of Nearly Everything: Special Illustrated Edition. Toronto: Doubleday Canada. ISBN 978-0-385-66198-0.
- Khanna, Dev Raj (2004). Human Evolution. Discovery Publishing House. p. 195. ISBN 978-8171417759. Retrieved 30 March 2013.
African H. erectus, with a mean stature of 170 cm, would be in the tallest 17 percent of modern populations, even if we make comparisons only with males
- Roylance, Frank D. Roylance (6 February 1994). "A Kid Tall For His Age". Baltimore Sun. Retrieved 30 March 2013.
Clearly this population of early people were tall, and fit. Their long bones were very strong. We believe their activity level was much higher than we can imagine today. We can hardly find Olympic athletes with the stature of these people
- Roach, & Richmond. (2015). "Clavicle length, throwing performance and the reconstruction of the Homo erectus shoulder". Journal of Human Evolution, 80(C), 107–113.
- Beck, Roger B.; Black, Linda; Krieger, Larry S.; Naylor, Phillip C.; Shabaka, Dahia Ibo (1999). World History: Patterns of Interaction. Evanston, IL: McDougal Littell. ISBN 978-0-395-87274-1.[page needed]
- Richards, M. P. (December 2002). "A brief review of the archaeological evidence for Palaeolithic and Neolithic subsistence". European Journal of Clinical Nutrition. 56 (12): 1270–1278. doi:10.1038/sj.ejcn.1601646. ISSN 1476-5640. PMID 12494313.
- Lepre, C. J.; Roche, H.; Kent, D. V.; et al. (2011). "An earlier origin for the Achuelian". Nature. 477 (7362): 82–85. doi:10.1038/nature10372. PMID 21886161.
- Gibbons, Ann (13 March 1998). "Paleoanthropology: Ancient Island Tools Suggest Homo erectus Was a Seafarer". Science. 279 (5357): 1635–1637. doi:10.1126/science.279.5357.1635.
- Oldest stone tool ever found in Turkey discovered by the University of Royal Holloway London and published in ScienceDaily on December 23, 2014
- Rincon, Paul (29 April 2004). "Early human fire skills revealed". BBC News. Retrieved 12 November 2007.
- Pollard, Elizabeth (2015). Worlds Together, Worlds Apart. New York: Norton. p. 13. ISBN 978-0-393-92207-3.
- Pringle, Heather (2 April 2012), "Quest for Fire Began Earlier Than Thought", ScienceNOW, archived from the original on 15 April 2013, retrieved 4 April 2012
- James, Steven R. (February 1989). "Hominid Use of Fire in the Lower and Middle Pleistocene: A Review of the Evidence" (PDF). Current Anthropology. 30 (1): 1–26. doi:10.1086/203705. Retrieved 14 January 2020.
- Callaway, Ewen (2014). "Homo erectus made world's oldest doodle 500,000 years ago". Nature News. doi:10.1038/nature.2014.16477.
- Brahic, Catherine (3 December 2014). "Shell 'art' made 300,000 years before humans evolved". New Scientist. Retrieved 29 September 2018.
- Joordens, Josephine C.A.; et al. (2015). "Homo Erectus at Trinil on Java Used Shells for Tool Production and Engraving". Nature. 518: 228–231. doi:10.1038/nature13962. PMID 25470048.
- Henshilwood, C.S.; d'Errico, F.; Watts, I. (2009). "Engraved ochres from the Middle Stone Age levels at Blombos Cave, South Africa". J.Hum. Evol. 57: 27–47. doi:10.1016/j.jhevol.2009.01.005. PMID 19487016.
- d'Errico, F.; Garcıa Moreno, R.; Rifkin, R.F. (2012). "Technological, elemental and colorimetric analysis of an engraved ochre fragment from the Middle Stone Age levels of Klasies River Cave 1, South Africa". J.Archaeol. Sci. 39: 942–952. doi:10.1016/j.jas.2011.10.032.
- Dubois, E. Pithecanthropus Erectus, Eine Menschena ¨hnliche U ¨bergangsformaus Java (Landesdruckerei, 1894).
- Ruhlen, Merritt (1994). The origin of language: tracing the evolution of the mother tongue. New York: Wiley. ISBN 978-0-471-58426-1.
- Bower, Bruce (3 May 2006). "Evolutionary back story: Thoroughly modern spine supported human ancestor". Science News. 169 (18): 275–276. doi:10.2307/4019325. JSTOR 4019325.
- Leakey, Richard (1992). Origins Reconsidered. Anchor. pp. 257–258. ISBN 978-0-385-41264-3.
- Boehm, Christopher (1999). Hierarchy in the forest: the evolution of egalitarian behavior. Cambridge: Harvard University Press. p. 198. ISBN 978-0-674-39031-7.
- Zanolli, Clément, et al. “Inner Tooth Morphology of Homo Erectus from Zhoukoudian. New Evidence from an Old Collection Housed at Uppsala University, Sweden.” Journal of Human Evolution, vol. 116, Mar. 2018, pp. 1–13.
- Copes, Lynn E., and William H. Kimbel. “Cranial Vault Thickness in Primates: Homo Erectus Does Not Have Uniquely Thick Vault Bones.” Journal of Human Evolution, vol. 90, Jan. 2016, pp. 120–134.
- Delson E, Harvati K, Reddy D, et al. (April 2001). "The Sambungmacan 3 Homo erectus calvaria: a comparative morphometric and morphological analysis". The Anatomical Record. 262 (4): 380–397. doi:10.1002/ar.1048. PMID 11275970.
- Ciochon R, Long VT, Larick R, et al. (April 1996). "Dated co-occurrence of Homo erectus and Gigantopithecus from Tham Khuyen Cave, Vietnam". Proceedings of the National Academy of Sciences of the United States of America. 93 (7): 3016–3020. Bibcode:1996PNAS...93.3016C. doi:10.1073/pnas.93.7.3016. PMC 39753. PMID 8610161.
- Schuster, Angela M.H. (September–October 1998). "New Skull from Eritrea". Archaeology. Retrieved 3 October 2015.
- Kappelman J, Alçiçek MC, Kazanci N, Schultz M, Ozkul M, Sen S (January 2008). "First Homo erectus from Turkey and implications for migrations into temperate Eurasia". American Journal of Physical Anthropology. 135 (1): 110–116. doi:10.1002/ajpa.20739. PMID 18067194.
|Wikimedia Commons has media related to Homo erectus.|
- Homo erectus Origins – Exploring the Fossil Record – Bradshaw Foundation
- Archaeology Info
- Homo erectus – The Smithsonian Institution's Human Origins Program
- Possible co-existence with Homo Habilis – BBC News
- John Hawks's discussion of the Kocabas fossil
- Peter Brown's Australian and Asian Palaeoanthropology
- The Age of Homo erectus – Interactive Map of the Journey of Homo erectus out of Africa
- Human Timeline (Interactive) – Smithsonian, National Museum of Natural History (August 2016).