Harpactognathus (meaning "seizing/grasping jaw") was a genus of pterosaur found in the Late Jurassic-age Morrison Formation of Albany County, Wyoming, United States. It is based on NAMAL 101, a partial skull consisting of the snout, recovered from near Bone Cabin Quarry in 1996. The species name honors the discoverer, Joe Gentry, a volunteer for the Western Paleontological Laboratories, in Lehi, Utah.
Temporal range: Late Jurassic
Carpenter, Unwin, Cloward, Clifford Miles, and Clark Miles, 2003
Its describers found it to be most similar to Scaphognathus among pterosaurs, albeit substantially larger (estimated skull length of 280–300 mm (11–12 in), estimated wingspan of at least 2.5 m (8.2 ft)). Because of the similarity, Harpactognathus was assigned to the subfamily Scaphognathinae of the family Rhamphorhynchidae. This genus is also notable for having a low bony crest running all the way to the tip of the beak (pterosaur bone crests usually do not reach the tip) and for being the oldest known Morrison Formation pterosaur, having been found in the Salt Wash Member (Kimmeridgian).
The authors hypothesized that scaphognathines specialized as aerial predators in inland freshwater habitats. However, more recent publications have suggested scaphognathines lacked speciations towards piscivory, and were likely terrestrial predators of small vertebrates or corvid-like generalists.
- Carpenter, K., Unwin, D.M., Cloward, K., Miles, C.A., and Miles, C. (2003). A new scaphognathine pterosaur from the Upper Jurassic Formation of Wyoming, USA. In: Buffetaut, E., and Mazin, J.- M. (eds.). Evolution and Palaeobiology of Pterosaurs. Geological Society of London, Special Publications 217:45-54.
- Witton, Mark (2013). Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press. p. 51. ISBN 978-0691150611.