The Coimbra Formation (Also known as Camadas de Coimbra or Calcários de S. Miguel) is a geological formation of Sinemurian (Lower Jurassic) age in the Lusitanian Basin of Portugal.[1] The unit represents a series of peritidal to intertidal facies of a Carbonate platform mostly of Obtusum–Oxynotum age, that gradually evolve to open marine/hemipelagic units Vale das Fontes Formation and Lemede formation.[2][3] This unit is known for its fossil content, including Invertertebrate and vertebrate ichnofossils and fossils.[4][5]

Coimbra Formation
Stratigraphic range: Early to Late Sinemurian 199–188 Ma
S. Pedro de Muel Beach, the main outcrop
TypeGeological formation
Underlies
Overlies
AreaLusitanian Basin
Thickness120-150 m
Lithology
PrimaryLimestones
Location
LocationCoimbra Region
Coordinates39.8° N, 9.0° W
RegionLusitanian Basin
Country Portugal
Type section
Named forThe Village of Coimbra
Thickness at type section~120 m (390 ft)
Coimbra Formation is located in Portugal
Coimbra Formation
Coimbra Formation (Portugal)

Paleoenvironment

edit
 
Stratigraphic column of the Lusitanian Basin

The sediments in the Coimbra Formation represent the shallow sections of an inner carbonate platform, adjacent to the Paleozoic basement in the E.[6] They are rich in organic matter and are found alongside marl layers in a shallow marine setting. The organic material is divided into three groups (A, B, and C), showing how it is spread across different layers.[1] The middle section mainly consists of group B, with a lot of microbial mat particles and very few plant particles. This layer has the highest level of Total organic carbon (TOC), indicating it was deposited in a restricted and stagnant area with little water movement.[7] In the upper section, all three groups (A, B, and C) are present. The lower part of this section contains more plant debris and fewer marine particles, suggesting a stronger influence from land and more oxygen in the environment, leading to lower TOC levels.[6] As we move up, group B shows a reduced land influence. At the top, group A takes over, with more marine particles and fewer plant materials, suggesting a transition to a more open marine environment. Overall, the Coimbra Formation is made up of shallow marine deposits formed in a low-energy setting, with occasional stronger events like storms or floods.[8] The area was a calm, shallow marine environment with limited water flow. Most sediments were fine-grained, showing low-energy conditions and a semi-arid climate. Microbialites formed during periods of slow sediment buildup, indicating low water movement. Occasional storms brought better oxygenated water, supporting brief marine life bursts. Fossils suggest shifts between restricted and open marine environments due to minor sea-level changes. Over time, open-marine conditions increased, with more typical marine fossils reflecting a significant rise in sea level.[8] Individual Stromatolite mounds are clearly defined, either as separate structures lined up next to each other or as irregular, flat to wavy layers that merge at the base.[8] Palynology includes Classopollis (Cheirolepidiaceae Conifer) and Botryococcus as the dominant continental components, while marine palynomorphs include the genera Tasmanites and Cymathiosphaera (Prasinophyte algae) as well Zygnemataceae.[6] The recent discovery of Dinosaur tracks revelated the presence of supratidal environments located in a shallow, tropical coastal area, likely near the shore.[4]

Fossil Content

edit
Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Mollusca

edit
Genus Species Location Material Notes Images
Asteroceras[9]
  • A. spp.
  • Sao Pedro de Muel
Isolated shells An Ammonite of the family Asteroceratidae
 
Reconstruction
Epophioceroides[9]
  • E. apertus
  • E. spp.
  • Sao Pedro de Muel
Isolated shells An Ammonite of the family Asteroceratidae
Gagaticeras[1]
  • G. spp.
  • Sao Pedro de Muel
Isolated shells An Ammonite of the family Echioceratidae
Mactromya[1]
  • M. spp.
  • Sao Pedro de Muel
Isolated shells A Bivalve of the family Mactromyidae
Oxynoticeras[1]
  • O. spp.
  • Sao Pedro de Muel
Isolated shells An Ammonite of the family Oxynoticeratidae
Pholadomya[1]
  • P. spp.
  • Sao Pedro de Muel
Isolated shells A Bivalve of the family Pholadomyidae
 
Example of specimen
Ptycharietites[9]
  • P. asteroceroides
  • P. ("subgen. indet. B") muellense
  • P. (Ptycharietites) heterogenus
  • P. (Pompeckiocreas) cf. oncocephalus
  • P. (Ptycharietites) ptychogenos
  • P. spp.
  • Sao Pedro de Muel
Isolated shells An Ammonite of the family Asteroceratidae
Unicardium[1]
  • U. costae
  • Sao Pedro de Muel
Isolated shells A Bivalve of the family Mactromyidae

Crustacea

edit
Genus Species Location Material Notes Images
Donzocythere[10]
  • D. cf. D. convergeas
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Cythereinae
Ektyphocythere[10]
  • E. retia
  • E. lacunosa
  • E. sinemiurana
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Progonocytherinae
Klieana[10]
  • K.? coimbraensis
  • Praia da Concha
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Cythereinae
Klinglerella[10]
  • "K." roselinae
  • Praia da Concha
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Progonocytherinae
Ljubimovella[10]
  • L.? frequens
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Cythereinae. The Second most abundant genus
Lutkevichinella[10]
  • L. hortonae
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Limnocytheridae
Marslatourella[10]
  • M. aff. M. heitoufensis
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Cythereinae
Monoceratina[10]
  • M. sp.
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Bythocytheridae
Phraterfabanella[10]
  • P. boomeri
  • Praia da Concha
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Cythereinae. Also recorded on the Rotzo Formation
Pseudomacrocypris[10]
  • P. cf. P. subtriangularis
  • Praia da Concha
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Macrocyprididae
Tropacythere[10]
  • T. normaniae
  • Sao Pedro de Muel
Isolated Carapaces & Valves A marine/brackish ostracodan member of the family Progonocytherinae

Brachiopoda

edit
Genus Species Location Material Notes Images
Terebratula[1]
  • “T.” ribeiroi
  • Sao Pedro de Muel
Isolated shells A marine Branchiopod of the family Terebratulidae
 
Specimen of the genus
Zeilleria[1]
  • Z. ssp.
  • Sao Pedro de Muel
Isolated shells A marine Branchiopod of the family Zeilleriidae

Plesiosauria

edit
Genus Species Location Material Notes Images
Plesiopharos[5]
  • P. moelensis
  • Praia da Concha
ML2302 a partial skeleton belonging to a single individual A Plesiosaur, possible member of Plesiosauroidea
 
Plesiopharos reconstruction

Archosauria

edit
Genus Species Location Material Notes Images
Batrachopus[4]
  • B. isp.
  • Campo de Alvaiázere
Footprints Tracks referred to Crocodylomorpha, maybe marine Teleosauridae
Lusitanosaurus[11]
  • L. liasicus
  • Sao Pedro de Muel
Single partial left maxilla An Indeterminate Archosaur, previously thought to be a Thyreophoran dinosaur. It's exact provenance is unknow
Moyenisauropus[4]
  • M. lusitanicus
  • Campo de Alvaiázere
Footprints Tracks referred to the Dinosaurian group Thyreophora, probably from a taxon similar to Scelidosaurus.
 
Scelidosaurus, a possible relative of the Moyenisauropus trackmaker

See also

edit

References

edit
  1. ^ a b c d e f g h i Duarte, Luís Vítor; Silva, Ricardo Louro; Azerêdo, Ana Cristina; Comas-Rengifo, María José; Mendonça Filho, João Graciano (2023-01-13). "Shallow-water carbonates of the Coimbra Formation, Lusitanian Basin (Portugal): contributions to the integrated stratigraphic analysis of the Sinemurian sedimentary successions in the western Iberian Margin". Comptes Rendus. Géoscience. 354 (S3): 89–106. doi:10.5802/crgeos.144. ISSN 1778-7025.
  2. ^ Duarte, L. V.; Soares, A. F. (2002). "Litostratigrafia das séries margo-calcárias do Jurássico inferior da Bacia Lusitânica (Portugal)" (PDF). Com. Instituto Geológico e Mineiro. 89 (1): 135–154.
  3. ^ Duarte, L. V.; Silva, R. F. L.; Oliveira, L. V.; Rengifo, M. J. C.; Silva, F. (2010). "Organic-rich facies in the Sinemurian and Pliensbachian of the Lusitanian Basin, Portugal: Total Organic Carbon distribution and relation to transgressive-regressive facies cycles". Geologica Acta: 325–340.
  4. ^ a b c d Figueiredo, Silvério D.; de Carvalho, Carlos Neto; Cunha, Pedro P.; Duarte, Luís V.; Fonseca, Alexandre; Monteiro, Cláudio; Forte, João (2023-09-20). "The first dinosaurs in Iberia: a new dinosaur tracksite from the Sinemurian (Lower Jurassic) of Portugal". Historical Biology: 1–14. doi:10.1080/08912963.2023.2256751. ISSN 0891-2963.
  5. ^ a b Puértolas-Pascual, Eduardo; Marx, Miguel; Mateus, Octávio; Saleiro, André; Fernandes, Alexandra; Marinheiro, João; Tomás, Carla; Mateus, Simão (2021). "A new plesiosaur from the Lower Jurassic of Portugal and the early radiation of Plesiosauroidea". Acta Palaeontologica Polonica. 66. doi:10.4202/app.00815.2020. hdl:10362/123694. ISSN 0567-7920.
  6. ^ a b c Poças Ribeiro, N.; Mendonça Filho, J.G.; Duarte, L.V.; Silva, R.L.; Mendonça, J.O.; Silva, T.F. (2013). "Palynofacies and organic geochemistry of the Sinemurian carbonate deposits in the western Lusitanian Basin (Portugal): Coimbra and Água de Madeiros formations". International Journal of Coal Geology. 111: 37–52. doi:10.1016/j.coal.2012.12.006. ISSN 0166-5162.
  7. ^ Correia, G. G.; Duarte, L. V.; Pereira, A.; Silva, R. L. (2013-02-15). "Outcrop gamma-ray spectrometry: Applications to the Sinemurian–Pliensbachian organic-rich facies of the Lusitanian Basin (Portugal)". Journal of Iberian Geology. 38 (2). doi:10.5209/rev_jige.2012.v38.n2.40464. ISSN 1886-7995.
  8. ^ a b c Azerêdo, Ana C.; Silva, Ricardo L.; Duarte, Luís V.; Cabral, M. Cristina (2009-09-11). "Subtidal stromatolites from the Sinemurian of the Lusitanian Basin (Portugal)". Facies. 56 (2): 211–230. doi:10.1007/s10347-009-0198-0. ISSN 0172-9179.
  9. ^ a b c Dommergues, J.L.; Meister, C.; Rocha, R. B. (2010). "The Sinemurian ammonites of the Lusitanian Basin (Portugal): an example of complex endemic evolution". Palaeodiversity. 3 (59–87).
  10. ^ a b c d e f g h i j k Cabral, M. Cristina; Colin, Jean-Paul; Azeredo, Ana C.; Silva, Ricardo L.; Duarte, Luis V. (2015). "Brackish and marine ostracode assemblages from the Sinemurian of western Portugal, with descriptions of new species". Micropaleontology. 61 (1–2): 3–24. doi:10.47894/mpal.61.1.02. ISSN 0026-2803.
  11. ^ Lapparent, A. F. d.; Zbyszewski, G. (1951). "Un Stégosaurien nouveau dans le Lias du Portugal [A new stegosaurian in the Lias of Portugal]". Boletim do Museu e Laboratório Mineralógico e Geológico, Faculdade de Ciências da Universidade de Lisboa. 19 (6): 107–108.