This is a list of fossils found at Maotianshan Shales, whose most famous assemblage of organisms are referred to as the Chengjiang biota.[1]

The Maotianshan Shales are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation,[2] famous for their Konservat Lagerstätten, deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada.[1]

Ecdysozoa edit

Genus Species Material Notes Images

Acosmia

  • A. maotiania
  • 9 nearly complete specimens

A small ecdysozoan worm that was originally thought to have been a priapulid worm.

 
Fossil of Acosmia

Laojieella

  • L. thecata
  • One complete and one incomplete specimen

An ecdysozoan worm considered close to the priapulids, it has a spiny proboscis, an elliptical theca on the posterior part of the trunk, and a caudal appendage

Scalidophora edit

Palaeoscolecidomorpha edit

Genus Species Material Notes Images

Cricocosmia

  • C. jinningensis

A genus of palaeoscolecid worm. Fossils have been found with smaller worms seemingly attached to them.

 
Reconstruction of Cricocosmia with parasitic Inquicus

Mafangscolex

  • M. sinensis
  • M. yunnanensis

Maotianshania

  • M. cylindrica

An extinct worm-like creature that might be an early member of the nematomorpha.

 
Fossil of Maotianshania

Tabelliscolex

  • T. chengjiangensis
  • T. hexagonus
  • T. maanshanensis

A genus of palaeoscolecid worm

Tylotites

  • T. petiolaris

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Priapulida edit

Genus Species Material Notes Images

Corynetis

  • C. brevis
  • C. fortis

An archaeopriapulid worm known from the biota.

 
Reconstruction of Corynetis

Eximipriapulus

  • E. globocaudatus

A carnivorous priapulid-like worm that has been found living in the discarded shells of hyoliths.

 
Reconstruction of Eximipriapulus

Paraselkirkia

  • P. sinica

An archaeopriapulid worm that resembles another worm, Selkirkia, from the Burgess shale

 
Reconstruction of Paraselkirkia

Paratubiluchus

  • P. bicaudatus
 
Reconstruction of Paratubiluchus

Palaeopriapulites

  • P. parvus
 
Reconstruction of Palaeopriapulites
  • S. elongata

A burrowing archaeopriapulid worm also known from the burgess shale.

 
Fossil of Selkirkia columbia

Sicyophorus

  • S. rarus

A genus of archaeopriapulid worm.

 
Reconstruction of Sicyophorus

Xiaoheiqingella

  • X. peculiaris

A priapulid worm thought to belong to the crown group.

 
Reconstruction of Xiaoheiqingella

Panarthropoda edit

Lobopodia edit

Genus Species Material Notes Images

Antennacanthopodia

  • A. gracilis
  • Known from two specimens, making the species very rare in the Chengjiang.

An unarmoured lobopodian, apparently closely related to Onychophora. The long, paired, cirriform structures at the end of the body could have been sensory, or perhaps represent viscera not externally present in life.

 
Reconstruction of Antennacanthopodia

Cardiodictyon

  • C. catenulum

A highly elongate armoured lobopodian, with saddle-shaped sclerites and a lightly sclerotized head.

 
Reconstruction of Cardiodictyon

Diania

  • D. cactiformis

A heavily armoured lobopodian.

 
Reconstruction of Diania

Lenisambulatrix

  • L. humboldti
  • A single specimen.

A simple, unarmoured lobopodian.

 
Reconstruction of Lenisambulatrix

Onychodictyon

  • O. ferox
  • O. gracilis
  • Several complete specimens.

An enigmatic genus of lobopodian, potentially related to either the Luolishaniidae or to Tardigrada. Not always monophyletic.

 
Reconstruction of Onychodictyon ferox

Paucipodia

  • P. inermis
  • Known from well-preserved body fossils.

A large unarmoured lobopodian, commonly found in association with mass death-beds of legless worms and Eldonia. Thought to have been a scavenger.

 
Reconstruction of Paucipodia

Eoconchariidae edit

Genus Species Material Notes Images

Microdictyon

  • M. sinicum
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Fossil of Microdictyon sinicum

Luolishaniidae edit

Genus Species Material Notes Images

Luolishania

  • L. longicruris

Type genus of Luolishaniidae. The species Miraluolishania haikouensis is considered a juvenile synonym of Luolishania.

 
Reconstruction of Luolishania

Facivermis

  • F. yunnanicus

Derived Luolishaniid lobopodian with five setiferous sieving appednages, and lacking any walking limbs. Sometimes discovered living in constructed tubes, buried into the sediment.

 
Reconstruction of Facivermis

Hallucigenia

  • H. fortis

A Luolishaniid lobopodian with a bulbous sclerotized head and two pairs of small feeding appendages, borne on a short neck.

 
Reconstruction of Hallucigenia fortis

Siberiidae edit

Genus Species Material Notes Images

Megadictyon

  • M. haikouensis
  • Posterior-most section of the trunk is unknown.

A poorly known siberiid.

 
Reconstruction of Megadictyon cf. haikouensis

Jianshanopodia

  • J. decorata
  • Known from multiple large individuals, all from a singular exposure of the Chengjiang.

A giant siberiid lobopodian related to Megadictyon.

 
Reconstruction of Jianshanopodia

Gilled Lobopodians edit

Genus Species Material Notes Images

Omnidens

  • O. amplus
  • Numerous well preserved oral cones.

A possible giant gilled lobopodian, closely related to Pambdelurion. Also present in the Xiaoshiba Biota.

 
Diagrammatic reconstruction of Omnidens

Radiodonta edit

Radiodonta are a group of highly successful panarthropods. The Chengjiang Fauna includes a large number of Radiodont species, primarily from the clade Amplectobeluidae. While hurdiids are poorly known from the formation and surrounding region, several specimens attributable to the family have been discovered, but remain unnamed.

Genus Species Material Notes Images

Houcaris

  • H. saron
  • Isolated appendages.

Previously described as "Anomalocaris" saron.

 
Appendage of Houcaris saron
Innovatiocaris
  • I. maotianshanensis
  • I. multispiniformis
  • Complete fossil and isolated appendages
Previously assigned to the Anomalocaris genus, and one of the most completely known radiodonts.
 
Restoration of I. maotianshanensis

Laminacaris

  • L. chimera
  • Giant isolated appendages, up to nearly 30 centimetres long.

A giant radiodont combining features from different clades.

 
Appendage of Laminacaris

Amplectobeluidae edit

Radiodonta are a group of highly successful panarthropods. The Chengjiang Fauna includes a large number of Radiodont species, primarily from the clade Amplectobeluidae. While hurdiids are poorly known from the formation and surrounding region, several specimens attributable to the family have been discovered, but remain unnamed.

Genus Species Material Notes Images

Amplectobelua

  • A. symbrachiata
  • Dozens of isolated appendages, and at least one complete body of a juvenile.

A large Amplectobeluid radiodont with long tail furcae. The frontal appendages formed a distinctive claw-like shape.

 
Reconstruction of Amplectobelua

Lyrarapax

  • L. trilobus
  • L. unguispinus
  • Known from appendages and complete body fossils, including neural anatomy.

Small Amplectobeluid radiodont. Some specimens of this genus represent some of the smallest known radiodonts.

 
Reconstruction of Lyrarapax unguispinus (left) and Lyrarapax trilobus (right)

Ramskoeldia

  • R. platyacantha
  • R. consimilis
  • Known from appendages, often associated with GLS plates, sclerites, and swimming flaps.

Large Amplectobeluid radiodonts, with crushing gnathobases.

 
Appendages of Ramskoeldia

Anomalocarididae edit

Radiodonta are a group of highly successful panarthropods. The Chengjiang Fauna includes a large number of Radiodont species, primarily from the clade Amplectobeluidae. While hurdiids are poorly known from the formation and surrounding region, several specimens attributable to the family have been discovered, but remain unnamed.

Genus Species Material Notes Images

Lenisicaris

  • L. lupata
  • Isolated appendages.

A second species is included in the genus, Lenisicaris pennsylvanica, from the Kinzers Formation.

 
Appendage of Lenisicaris

Arthropoda edit

Genus Species Material Notes Images
Chuandianella
  • C. ovata
  • Several body fossils and carapaces
An enigmatc bivalved arthropod originally considered close to Waptia. It was then discovered that it lacked mandibles and therefore does not belong to Hymenocarina, the Waptiidae family, and even the larger mandibulata clade.
 
Reconstruction of Chuandianella

Combinivalvula

  • C. chengjiangensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

An enigmatic bivalved arthropod.

Cucumericrus

  • C. decorata

Multiple Specimens.

An enigmatic arthropod that has been tentatively classified as a possible radiodont.

 
Reconstructed trunk appendage of Cucumericrus

Cyathocephalus

  • C. bispinosus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Dianchia

  • D. mirabilis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Diplopyge

  • D. forcipatus
  • D. minutus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Dongshanocaris

  • D. foliiformis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod that is currently "too poorly preserved to verify their identity as a valid taxa".

Erratus

  • E. sperare
  • several carapaces
  • eyes and other body parts

A bivalved arthropod that shows the origins of lungs and legs in early arthropods.

 
Restoration of E. sperare

Ercaia

  • E. miniscula
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

An arthropod that has been speculated to be an early crustacean.

Forfexicaris

  • F. valida
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small bivalved arthropod that somewhat resembles ostracods, but also has large, upward facing appendages with spines.

 
Reconstruction of Forfexicaris

Glossocaris

  • G. occulatus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Isoxys

  • I. auritus
  • I. curvirostratus
  • I. paradoxus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A basal nektonic arthropod that possessed a bivalved carapace, and large circular eyes.

 
Restoration of Isoxys curvirostratus

Jianshania

  • J. furcatus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small, somewhat enigmatic arthropod known from the biota

 
Fossil of Jianshania

Kunmingocaris

  • K. bispinosus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Kylinxia

  • K. zhangi
  • almost complete fossil
  • Isolated frontal appendages

An early arthropod that may show the "missing link" between early stem-arthropods, and more advanced true arthropods. It possessed frontal appendages similar to radiodonts, and an eye arrangement similar to the opabiniids.

 
Reconstruction of Kylinxia

Mafangia

  • M. subscalaria
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Mafangocaris

  • M. multinodus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Occacaris

  • O. oviformis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A predatory arthropod that had spiked frontal appendages similar to those of the megacheirans, and was previously thought to have belonged to that grouping

 
Reconstruction of Occacaris

Ovalicephalis

  • O. mirabilis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Parapaleomerus

  • P. sinensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

An arachnomorph arthropod of the group strabopida. Unlike the other members of strabopida, Parapaleomerus lacks dorsal eyes and only possesses ten trunk tergites. The telson has been described as trapezoidal in shape.

 
Reconstructed carapace of Parapaleomerus

Pisinnocaris

  • P. subconigera
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Pseudoiulia

  • P. cambriensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A poorly known arthropod that, in 2013, was suggested to be a member of the family Kootenichelidae, alongside Kootenichela and Worthenella.

Pterotum

  • P. triacanthus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Rhombicalvaria

  • R. acanthi
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Sunella

  • S. grandis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Syrrhaptis

  • S. intes
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Tanglangia

  • T. caudata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Tauricornicaris

  • T. latizonae
  • T. oxygonae
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A giant arthropod once misidentified as radiodont.

Trigoides

  • T. aclis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Urokodia

  • U. aequalis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

An arthropod that had segmentation which resembled that of a millipede, as well as head and tail shields with thorny spikes. Recently, it was grouped in the mollisoniid order alongside Mollisonia, Thelxiope, and Corcorania, making it a stem-chelicerate.

 
Diagrammatic reconstruction of Urokodia

Wutingella

  • W. bindosa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Yiliangocaris

  • Y. ellipticus
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Yunnanocaris

  • Y. megista
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Zhenghecaris

  • Z. shankouensis
  • sclerites

Large-sized arthropod carapace, originally described as bivalved arthropod close to Thylacocephala, later reinterpreted as hurdiid radiodont close to Cambroraster, its classification is still discussed.

 
Fossil of Zhenghecaris

Hymenocarina edit

Genus Species Material Notes Images

Branchiocaris

  • B. yunnanensis
  • Several isolated carapaces

Also known in the Burgess Shale by B. pretiosa, B. yunnanensis is only known from isolated carapaces, although it is thought that this may be because in this species the carapace covered the entire body. It also has two morphotypes.

 
Reconstruction of the related Branchiocaris pretiosa

Canadaspis

  • C. laevigata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A hymenocarine arthropod, also known from the Burgess Shale by C. perfecta, although some authors believe that both species should be separated into different genera. They were mainly benthic animals.

 
Reconstruction of Canadaspis laevigata

Clypecaris

  • C. pteroidea
  • C. serrata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A hymenocarine arthropod. The species in the genus are primarily distinguished by the presence of a serrated edge on the front of the carapace of C. serrata. C. serrata is noted for the modification of an anterior pair of limbs into spined grasping appendages, indicating a predatory lifestyle.

 
Reconstruction of Clypecaris serrata

Ercaicunia

  • E. multinodosa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A hymenocarine arthropod that had a bivalved carapace which covered about a third of its total body-length, and had up to six serrations on its forward edge. The head had a pair of large uniramous antennae, as well as a smaller pair of secondary antennae, and a pair of mandibles and maxillae.

 
Reconstruction of Ercaicunia

Erjiecaris

  • E. minusculo
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small euarthropod, and unlike other hymenocarines, it possibly had eyes directly on top of its carapace. It was most likely a nektobenthic animal.

 
Reconstruction of Erjiecaris

Pectocaris

  • P. spatiosa
  • P. eurypetala
  • P. inopinata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A bivalved arthropod of the hymenocarina grouping. The species P. spatiosa is regarded as a filter feeder, using the setae on the endites of their limbs to filter out matter from the water column

 
Reconstruction of Pectocaris spatiosa

Synophalos

  • S. xynos
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A hymenocarine notable for fossils showing a bizarre chain of multiple individuals. Although the purpose for this behavior is unknown, it may have served a reproductive, migratory, or defensive purpose, with the authors of the describing paper of this genus considering migration as its most likely function.

 
Reconstruction of a Synophalos chain
Tuzoia
  • T. sinensis
  • several carapaces and soft body parts
A large hymenocarine that possessed a semicircular bivalved carapace, and was one of the largest arthropods from the Cambrian
 
Life restoration of Tuzoia sp.
Jugatacaris A. agillis
  • several carapaces and soft body parts
A hymenocarine arthropod. The carapace of the animal was around 28 to 37 millimetres (1.1 to 1.5 in) in length, with a pronounced ridge at the top of the carapace separating the two valves, which formed a fin-like structure raised above the carapace. The head had a pair of stalked eyes, as well as a dumbbell shaped medial eye between them. The head also bore a pair of mandibles as well as at least one and possibly two pairs of antennules.
 
Reconstruction of Jugatacaris
Xiazhuangocaris X. chenggongensis
  • a single specimen (NIGP 172765), which only preserves the carapace and trunk regions
A hymenocarine with no known head fossils. The material known show that the carapace had a pronounced pair of notches at its front, as well as a posterior notch at its rear. The body had at least 13 tergite-pleurite rings, which terminate in a pair of rounded caudal rami, which are fringed with setae.
 
Reconstruction of Xiazhuangocaris

Fuxianhuiida edit

Genus Species Material Notes Images

Chengjiangocaris

  • C. longiformis
  • C. kunmingensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A Fuxianhuiid arthropod with numerous appendages. It is one of the most complete arthropods from the lower Cambrian.

 
Reconstruction of Chengjiangocaris

Fuxianhuia

  • F. protensa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A large arthropod named after Fuxian Lake. Some adult specimens are found closely associated with numerous juveniles, indicating a level of parental care.

 
Reconstruction of Fuxianhuia with associated juveniles

Bradoriida edit

Genus Species Material Notes Images

Comptaluta

  • C. inflata
  • C. kailiensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Indiana

  • I. sp.
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
CT-scan of a fossil of Indiana sp.

Jiucunella

  • J. paulula
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Kunmingella

  • K. angustacostata
  • K. douvillei
  • K. guanshanensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Restoration of Kunmingella douvillei

Kunyangella

  • K. cheni
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Liangshanella

  • L. liangshanensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Malongella

  • M. bituerculata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Parakunmingella

  • P. malongensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Spinokunmingella

  • S. typica
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Tsunyiella

  • T. daindongensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Megacheira edit

Genus Species Material Notes Images

Haikoucaris

  • H. ercaiensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Haikoucaris
Leanchoiliidae edit
Genus Species Material Notes Images

Alalcomenaeus

  • A. cambricus

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Alalcomenaeus

Leanchoilia

  • D. asiatica
  • L. illecebrosa

One of the several species of Leanchoilia, it has known juvenile specimens and the labrum morphology known.

 
Fossil of Leanchoilia illecebrosa
Jianfengiidae edit
Genus Species Material Notes Images

Fortiforceps

  • F. foliosa
 
Reconstruction of Fortiforceps
Sklerolibyon *S. maomima
 
Reconstruction of Sklerolibyon

Jianfengia

  • J. multisegmentalis
 
Reconstruction of Jianfengia

Parapeytoia

  • P. yunnanensis

A jianfengiid once misidentified as a radiodont

 
Reconstruction of known elements from Parapeytoia

Marrellomorpha edit

Genus Species Material Notes Images

Primicaris

  • P. larvaformis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Primicaris

Artiopoda edit

Genus Species Material Notes Images

Acanthomeridion

  • A. serratum

Multiple Specimens.

A sessile, marine annelid tube worm of the family Serpulidae. Its affinities with the genus Serpula are controversial, since the genus is known mostly since Cretaceous strata. Although there are other fossils assigned to the genus on same age deposits of France. Presumably these specimens have fallen from their growth areas.[3]

 
Reconstruction of the carapace of Acanthomeridion

Almenia

  • A. spinosa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A polychaete of the family Dorvilleidae inside Eunicida. It was considered as part of the genus Halla. Eunicidan species with prionognath jaws, absent on Bituminous layers

Cindarella

  • C. eucalla
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Cindarella

Kuamaia

  • K. lata
  • K. muricata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Kuamaia lata

Misszhouia

  • M. longicaudata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Fossil of Misszhouia

Naraoia

  • N. spinosa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Naraoia spinosa

Panlongia

  • P. spinosa
  • P. tetranudosa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Fossil of Panlognia

Pygmaclypeatus

  • P. daiensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
CT-scan of a specimen of Pygmaclypeatus

Retifacies

  • R. abnormalis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Retifacies

Saperion

  • S. glumaceum
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Sidneyia

  • S. sinica
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstructed model of Sidneyia

Sinoburius

  • S. lunaris
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Sinoburius lunaris

Skioldia

  • S. aldna
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Squamacula

  • S. clypeata
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Squamacula

Xandarella

  • X. spectaculum
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Xandarella

Trilobita edit

Genus Species Material Notes Images

Eoredlichia

  • E. intermedia
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Reconstruction of Eoredlichia

Kuanyangia

  • K. pustulosa
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Palaeolenus

  • E. miniscula
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Fossil pair of Palaeolenus

Tsunyidiscus

  • T. aclis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Line drawing of Tsunyidiscus niutitangensis

Wutingapsis

  • W. tingi
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

Yunnanocephalus

  • Y. yunnanensis
  • Single Scolecodont
  • Incomplete Indeterminate Scolecodont

A small arthropod related to Leanchoilia, also known from the Burgess Shale.

 
Fossil of Yunnanocephalus

Spiralia edit

Chaetognatha edit

Genus Species Material Notes Images

Eognathacantha

  • E. ercainella
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Eognathacantha

Lophotrochozoa edit

Mollusca edit

Genus Species Material Notes Images

Orthrozanclus

  • O. elongata
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Orthrozanclus elongata

Petalilium

  • P. latus
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Fossil of Nectocaris pteryx

Wiwaxia

  • W. papillo
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of a juvenile Wiwaxia corrugata, which resembles W. papillo in its apparent lack of dorsal sclerite spines.

Hyolitha edit

Genus Species Material Notes Images

Ambrolinevitus

  • A. maximus
  • A. platypluteus
  • A. ventricosus
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Burithes

  • B. yunnanensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Glossolithes

  • G. magnus
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Linevitus

  • L. billingsi
  • L. flabellaris
  • L. opimus
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Phoronida edit

Genus Species Material Notes Images

Iotuba

  • I. chengjiangensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Fossil of Iotuba chengjiangensis

Eophonoris

  • E. chengjiangensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Entoprocta edit

Genus Species Material Notes Images

Cotyledion

  • C. tylodes
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Cotyledion

Cornulitida edit

Genus Species Material Notes Images

Cambrocornulitus

  • C. rarus
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Brachiopoda edit

Genus Species Material Notes Images

Diandongia

  • D. pista
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Heliomedusa

  • H. orienta
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Fossil of Heliomedusa orienta

Lingulella

  • H. orienta
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Fossil of Lingulella caelata

Lingulellotreta

  • L. malongensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Longtancunella

  • L. chengjiangensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Deuterostomia edit

Genus Species Material Notes Images

Yunnanozoon

  • Y. lividium
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Fossil of Yunnanozoon

Chordata edit

Genus Species Material Notes Images

Cathaymyrus

  • C. diadexus
  • C. haikouensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of C. diadexus and Vetulocystis

Haikouichthys

  • H. ercaicunensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Haikouichthys

Myllokunmingia

  • M. fengjiaoa
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Myllokunmingia

Zhongjianichthys

  • Z. rostratus
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Zhongjianichthys

Zhongxiniscus

  • Z. intermedius
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Zhongxiniscus

Tunicata edit

Genus Species Material Notes Images

Shankouclava

  • S. anningense
  • S. shankouense
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Shankouclava anningense

Vetulicolia edit

Genus Species Material Notes Images

Beidazoon

  • B. venustum
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Beidazoon

Didazoon

  • D. haoae
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Beidazoon

Pomatrum

  • P. ventralis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Beidazoon

Vetulicolia

  • V. cuneata
  • V. gantoucunensis
  • V. rectangulata
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Vetulicola rectangulata and V. cuneata

Yuyuanozoon

  • Y. magnificissimi
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Yuyuanozoon

Ambulacraria edit

Echinodermata edit

Genus Species Material Notes Images

Dianchicystis

  • D. jianshanensis
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

Vetulocystis

  • V. catenata
  • Dozens of complete specimens.

Microdictyon is known from slcerite microfossils around the globe, while fossils from the Chengjiang are the only in the world to preserve the soft anatomy.

 
Reconstruction of Vetulocystis (bottom)


Phylum Ctenophora edit

Genus Species Material Notes Images

Maotianoascus

  • M. octanarius
 
Reconstruction of Maotianoascus octanarius

Sinoascus

  • S. paillatus
 
Reconstruction of Sinoascus papillatus
Stromatoveris
  • S. psygmoglena
 
Reconstruction of Stromatoveris psygmoglena

Phylum Porifera edit

15 species

Enigmatic edit

24 species

 
Phlogites longus

Kingdom Protista (algae) edit

Cnidaria edit

3 species

References edit

  1. ^ a b Fossils of the Chengjiang Maotianshan Shale
  2. ^ Lipps, J. H.; Signor, P. W (1992). Origin and early evolution of the Metazoa. ISBN 978-0-306-44067-0.
  3. ^ Cite error: The named reference BlackFo was invoked but never defined (see the help page).

Maotianshan Category:Maotianshan shales fossils