Turkestan short-toed lark

The Turkestan short-toed lark (Alaudala heinei) is a species of lark in the family Alaudidae. It is found in Ukraine and central Turkey through parts of Central Asia and southern Siberia west to south-central Mongolia and south to southern Afghanistan.[1] This species and the Mediterranean short-toed lark (A. rufescens) were formerly considered conspecific and called the lesser short-toed lark, but a 2020 study recovered them as distinct species.[2][3]

Turkestan short-toed lark
Naturalis Biodiversity Center - RMNH.AVES.31393 1 - Calandrella rufescens heinei (Homeyer, 1873) - Alaudidae - bird skin specimen.jpeg
Specimen at Naturalis Biodiversity Center
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Passeriformes
Family: Alaudidae
Genus: Alaudala
A. heinei
Binomial name
Alaudala heinei
(Homeyer, 1873)


Four subspecies are recognized:[1]

  • A. h. pseudobaetica - (Stegmann, 1932): Found in eastern Turkey, Transcaucasia and northern Iran
  • A. h. heinei - (Homeyer, 1873): Found from Ukraine to eastern Kazakhstan
  • A. h. aharonii - (Hartert, 1910): Found in central Turkey
  • A. h. persica - Sharpe, 1890: Found in eastern and southern Iraq to southern and south-western Afghanistan


  1. ^ a b "Nicators, reedling & larks « IOC World Bird List". www.worldbirdnames.org. Retrieved 2021-01-19.
  2. ^ "Alaudala [rufescens or heini, excl. niethammeri] (Mediterranean or Turkestan Short-toed Lark [excl. niethammeri]) - Avibase". avibase.bsc-eoc.org. Retrieved 2021-01-30.
  3. ^ "Multiple species delimitation approaches applied to the avian lark genus Alaudala". Molecular Phylogenetics and Evolution. 154: 106994. 2021-01-01. doi:10.1016/j.ympev.2020.106994. ISSN 1055-7903.