Melanella

Melanella is a genus of very small ectoparasitic sea snails, marine gastropod molluscs or micromolluscs in the family Eulimidae.

Melanella
Melanella candida
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Caenogastropoda
Order:	Littorinimorpha
Family:	Eulimidae
Genus:	Melanella Bowdich, 1822
Synonyms
Balcis Leach, 1847 Euliamaustra Laseron, 1955 Eulima (Balcis) Eulimaustra Laseron, 1955 Melanella (Balcis) Melanella (Melanella) Bowdich, 1822

Species

The following species are recognised in the genus Melanella:

• Melanella abida Dall, 1927
• Melanella acanthyllis (R. B. Watson, 1883)
• Melanella acicula (A. Gould, 1849)
• Melanella aciculata (Pease, 1861)
• Melanella acuformis (G. Nevill & H. Nevill, 1875)
• Melanella adamantina (de Folin, 1867)
• Melanella adiastalta Souza, Pimenta & J. Barros, 2021
• Melanella aequatorialis (Thiele, 1925)
• Melanella aethiopica (Thiele, 1925)
• Melanella aethria (Melvill, 1918)
• Melanella agulhasensis (Thiele, 1925)
• Melanella alba (da Costa, 1778)
• Melanella alertae (Dell, 1956)
• Melanella alfredensis Bartsch, 1915
• Melanella algoensis (E. A. Smith)
• Melanella amabilis (Brazier, 1877)
• Melanella amblia (R. B. Watson, 1883)
• Melanella amblytera (Verrill & Bush, 1900)
• Melanella ameliae Engl, 2007
• Melanella anachorea Dall, 1927
• Melanella anapetes Souza, Pimenta & J. Barros, 2021
• Melanella antarctica (Strebel, 1908)
• Melanella archeyi (Finlay, 1928)
• †Melanella arenularia (Gougerot in Perreau, 1969)
• Melanella arleyi Espinosa, Ortea & Magaña, 2001
• Melanella atlantica (E. A. Smith, 1890)
• Melanella attenuata (G. B. Sowerby II, 1866)
• Melanella aucklandica (Suter, 1909)
• Melanella augur (Angas, 1865)
• †Melanella badenia (Laws, 1939)
• Melanella baldra Bartsch, 1917
• Melanella becki (Turton, 1932)
• Melanella bipartita (Mörch, 1859)
• Melanella borealis (Bartsch, 1917)
• Melanella boscheineni Engl, 2004
• Melanella boscii (Payraudeau, 1826)
• Melanella bovicornu (Pilsbry, 1905)
• Melanella breviuscula (Dunker, 1875)
• Melanella brunnimaculata (Kay, 1979)
• Melanella bryani Pilsbry, 1918
• Melanella callistemma Dall, 1927
• Melanella candida (Marrat, 1880)
• Melanella catalinensis Bartsch, 1917
• Melanella chariessa (Verrill, 1884)
• Melanella chydaea (Watson, 1883)
• Melanella cifara Bartsch, 1915
• Melanella clavula (A. Adams, 1861)
• †Melanella collator (Laws, 1944)
• Melanella compactilis (Locard, 1891)
• Melanella conoidalis (G. B. Sowerby II, 1865)
• Melanella conoidea (Kurtz & W. Stimpson, 1851)
• Melanella convexa (E. A. Smith, 1907)
• Melanella corintonis (Hertlein & A. M. Strong, 1951)
• Melanella cosmia Bartsch, 1917
• Melanella costellata (Dautzenberg & H. Fischer, 1897)
• Melanella cumingii (A. Adams, 1854)
• Melanella cunaeformis (May, 1915)
• Melanella cuspidata (A. Adams, 1854)
• Melanella cyclopia Scuderi, Tisselli & Micali, 2022
• Melanella cylindrica (Thiele, 1925)
• Melanella debilis (A. Adams, 1861)
• Melanella decagyra (Melvill & Standen, 1903)
• Melanella decipiens (Hedley, 1899)
• Melanella densicostata Bouchet & Warén, 1986
• †Melanella distorta (Defrance, 1823)
• Melanella doederleini (Brusina, 1886)
• Melanella draconis Bartsch, 1917
• †Melanella dubusi (Cossmann & Pissarro, 1902)
• Melanella dufresnii Bowdich, 1822
• Melanella eburnea (Megerle von Mühlfeld, 1824)
• Melanella elenensis (Bartsch, 1924)
• Melanella elsa Thiele, 1930
• Melanella eulimoides (C. B. Adams, 1850)
• Melanella exasperata (Preston, 1905)
• Melanella exilis (Pease, 1863)
• Melanella exulata (E. A. Smith, 1915)
• Melanella falcata (Carpenter, 1865)
• Melanella fasciata (R. B. Watson, 1883)
• Melanella flexuosa (A. Adams, 1854)
• Melanella frielei (H. K. Jordan, 1895)
• Melanella gagei Bouchet & Warén, 1986
• †Melanella geoffreyi (Laws, 1936)
• Melanella glypta Bouchet & Warén, 1986
• Melanella gracilis (C. B. Adams, 1850)
• Melanella gradata (Cotton & Godfrey, 1932)
• Melanella grandis (A. Adams, 1853)
• Melanella gratiosa (Thiele, 1925)
• Melanella grippi (Bartsch, 1917)
• Melanella guenteri Engl, 2004
• Melanella hadalis Knudsen, 1964
• Melanella halia Bartsch, 1917
• Melanella halorhaphe (Dautzenberg & H. Fischer, 1896)
• Melanella hastata (G. B. Sowerby I, 1834)
• Melanella hebes (R. B. Watson, 1883)
• Melanella hemphilli Bartsch, 1917
• Melanella hians (R. B. Watson, 1883)
• Melanella hypsela (Verrill & K. J. Bush, 1900)
• Melanella ima Pilsbry, 1918
• Melanella indica (Preston, 1905)
• Melanella insculpta Bouchet & Warén, 1986
• Melanella iotoides Bartsch, 1916
• Melanella ira Dall, 1927
• Melanella jeffreysii (Tryon, 1886)
• Melanella jucunda (Thiele, 1925)
• Melanella junieri Espinosa, Ortea & Magaña, 2006
• †Melanella kaiparaensis (Laws, 1939)
• Melanella kanaka Pilsbry, 1917
• Melanella karolinae Engl, 2006
• Melanella kermadecensis W. R. B. Oliver, 1915
• Melanella kuronamako (Habe, 1952)
• Melanella lactea (A. Adams, 1854)
• Melanella lastra Bartsch, 1917
• Melanella laurae (Friele, 1886)
• †Melanella lautoides Finlay & Marwick, 1937
• Melanella letsonae Pilsbry, 1917
• Melanella levantina (Oliverio, Buzzurro & R. Villa, 1994)
• Melanella lubrica (Monterosato, 1890)
• Melanella luchuana (Pilsbry, 1901)
• Melanella lucida (A. E. Verrill, 1884)
• Melanella luminosa B. A. Marshall, 1997
• Melanella lunata Pilsbry, 1918
• Melanella magnifica (Bartsch, 1915)
• Melanella major (G. B. Sowerby I, 1834)
• Melanella mamilla (May, 1915)
• Melanella maoria (Powell, 1940)
• Melanella martarum Souza, Asorey & Sellanes, 2024
• Melanella martinii (A. Adams, 1854)
• Melanella martynjordani (H. K. Jordan, 1895)
• Melanella marviva Espinosa, Ortea & Magaña, 2006
• Melanella maui (Dell, 1952)
• Melanella micans (P. P. Carpenter, 1865)
• Melanella microsculpta Bouchet & Warén, 1986
• Melanella mimus Pilsbry, 1918
• Melanella minor (Pallary, 1904)
• Melanella modesta (Thiele, 1925)
• Melanella modesta Thiele, 1930
• Melanella montereyensis Bartsch, 1917
• Melanella monterosatoi (Monterosato, 1890)
• Melanella muelleriae (Sturany, 1903)
• Melanella mundula (A. Adams, 1861)
• Melanella myriotrochi Bouchet & Warén, 1986
• Melanella nilae Engl, 2006
• Melanella nisonida (Melvill, 1906)
• Melanella obtusa (Laseron, 1955)
• Melanella obtusoapicata Bouchet & Warén, 1986
• Melanella ochsneri Bartsch, 1917
• Melanella odontoidea (A. Adams, 1861)
• Melanella ogasawarana (Pilsbry, 1905)
• Melanella oldroydae Bartsch, 1917
• Melanella oleacea (Kurtz & W. Stimpson, 1851)
• Melanella olssoni (Bartsch, 1924)
• Melanella opaca (G. B. Sowerby II, 1865)
• Melanella opalina (J. T. Marshall, 1901)
• Melanella ophiodon Dall, 1927
• Melanella orientalis Agamennone, Micali & Siragusa, 2020
• Melanella orphanensis A. H. Clarke, 1974
• Melanella orthophyes (Sturany, 1903)
• Melanella otagoensis (Powell, 1927)
• Melanella oxyacme (Suter, 1908)
• Melanella oxytata (R. B. Watson, 1883)
• Melanella pandata (A. Adams, 1861)
• Melanella paraabida Souza, Pimenta & J. Barros, 2021
• Melanella parallela Dall, 1927
• †Melanella parviapertura Landau, La Perna & Marquet, 2006
• †Melanella patutahiensis Marwick, 1931
• Melanella paxillus (Hedley, 1904)
• Melanella peninsularis Bartsch, 1917
• Melanella penna Dall, 1927
• †Melanella pepediazi Landau, La Perna & Marquet, 2006
• Melanella perplexa W. R. B. Oliver, 1915
• Melanella persimilis (Kuroda & Habe, 1971)
• Melanella petitiana (Brusina, 1869)
• Melanella pisinna Rehder, 1980
• Melanella planisutis Bouchet & Warén, 1986
• †Melanella plioiberica Landau, La Perna & Marquet, 2006
• Melanella polita (Linnaeus, 1758)
• †Melanella polygyra (Charlesworth, 1851)
• †Melanella pontilis P. A. Maxwell, 1992
• Melanella praecurta (Pallary, 1904)
• Melanella producta (P. P. Carpenter, 1863)
• Melanella pseudoglabra (Dautzenberg & H. Fischer, 1912)
• Melanella puhana (Dell, 1956)
• Melanella reticulata Bouchet & Warén, 1986
• Melanella retrorsa (G. B. Sowerby II, 1865)
• †Melanella retusa (Pezant, 1911)
• Melanella richardi (Dautzenberg & H. Fischer, 1896)
• Melanella robusta (A. Adams, 1861)
• Melanella roegerae (Cotton & Godfrey, 1932)
• Melanella salvadori Castellanos, Rolán & Bartolotta, 1987
• Melanella sankurieae Engl, 2004
• Melanella sansibarica (Thiele, 1925)
• Melanella sarissa (R. B. Watson, 1883)
• Melanella scarifata Gofas & Rueda, 2014
• †Melanella seguenzai Di Geronimo & La Perna, 1999
• Melanella shibana (Yokoyama, 1927)
• Melanella siberutensis (Thiele, 1925)
• Melanella similior Bouchet & Warén, 1986
• †Melanella similis (d'Orbigny, 1852)
• Melanella solida (G. B. Sowerby II, 1865)
• Melanella solidula (A. Adams & Reeve, 1850)
• Melanella soliduloides (Habe, 1951)
• Melanella solitaria (E. A. Smith, 1915)
• †Melanella sororcula (Wrigley, 1945)
• †Melanella sowerbyi (Newton, 1891)
• Melanella spina (O. Boettger, 1893)
• Melanella spinosa W. R. B. Oliver, 1915
• Melanella spiridioni (Dautzenberg & H. Fischer, 1896)
• Melanella stalioi (Brusina, 1869)
• Melanella stamina Dall, 1927
• Melanella subantarctica (Strebel, 1908)
• †Melanella subemarginata (Deshayes, 1862)
• Melanella subula (A. Adams, 1861)
• †Melanella sulculata (Wrigley, 1945)
• Melanella tanabensis Takano, Tanaka & Kano, 2019
• Melanella teinostoma (A. Adams, 1854)
• Melanella teramachii (Habe, 1952)
• Melanella thalia Bartsch, 1915
• Melanella thersites (P. P. Carpenter, 1864)
• Melanella tia (Bartsch, 1926)
• Melanella tortuosa (A. Adams & Reeve, 1850)
• Melanella townsendi Bartsch, 1917
• Melanella translucens (Monterosato, 1890)
• Melanella trunca (R. B. Watson, 1897)
• Melanella truncata (Suter, 1908)
• Melanella turbonilloides Bouchet & Warén, 1986
• Melanella turrita Bouchet & Warén, 1986
• †Melanella tutamoensis Marwick, 1931
• Melanella variabilis (Schepman, 1913)
• Melanella vegrandis (Murdoch & Suter, 1906)
• Melanella versa Dall, 1927
• †Melanella waiaotea (Laws, 1941)
• †Melanella waikomitica (Laws, 1939)
• †Melanella wareni Landau, La Perna & Marquet, 2006 
• Melanella yamazii (Habe, 1952)
• Melanella zugnigae Espinosa, Ortea & Magaña, 2001

Melanella martinii, abapertural view

Three shells of Melanella solidula

The Indo-Pacific Molluscan Database also includes the following species with names in current use

• Melanella australiensis (Thiele, 1930)
• Melanella campyla (Watson, 1883)
• Melanella carneola (Gould, 1861 in 1859–61)
• Melanella curvata (A. Adams, 1861-b)
• Melanella dentaliopsis (A. Adams, 1861-b)
• Melanella flexa (A. Adams, 1861-b)
• Melanella labiosa (Sowerby II, 1834-b)
• Melanella musta (Yokoyama, 1928)
• Melanella nivea (A. Adams, 1861): species inquirenda
• Melanella oblonga (Boettger)
• Melanella pinguicula (A. Adams, 1861)
• Melanella reclinata (A. Adams, 1861)
• Melanella scitula (A. Adams, 1861-b)
• Melanella semitorta (A. Adams, 1861-b)
• Melanella stenostoma (A. Adams, 1861-b)
• Melanella stylata (A. Adams, 1861-b)
• Melanella valida (A. Adams, 1861-b)
• Melanella venusta (Pease, 1868-b)

References

• Leach, 1847 [in Gray] Annals and Magazine of Natural History, 20: 271; Placed on the Official List by Opinion 1739, 1993, Bulletin of Zoological Nomenclature, 50(3): 242
• Warén A. (1984) A generic revision of the family Eulimidae (Gastropoda, Prosobranchia). Journal of Molluscan Studies suppl. 13: 1–96. page(s): 54
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213

External links

• Bowdich, T.E. 1822. Elements of Conchology, including the fossil genera and animals. Part 1. Univalves. Paris : Smith pp. i–xi, 14–73, index 1–3, 19 pls, 2 tables
• Gray, J. E. (1847). A list of the genera of recent Mollusca, their synonyma and types. Proceedings of the Zoological Society of London. (1847) 15: 129-219
• Laseron, C. 1955. Revision of the New South Wales eulimoid shells. The Australian Zoologist 12(2): 83–107, pls 1–3, figs 1–78 [85–86, 101] [Euliamaustra is an alternative original spelling (p. 85); spelling Eulimaustra selected by First Reviser action by McMichael & Whitley, 1961, The Marine Zoologist, 1(8): 124]
• Serge GOFAS, Ángel A. LUQUE, Joan Daniel OLIVER,José TEMPLADO & Alberto SERRA (2021) - The Mollusca of Galicia Bank (NE Atlantic Ocean); European Journal of Taxonomy 785: 1–114