Soricinae is a subfamily of small mammals in the shrew family Soricidae, which in turn is part of the order Eulipotyphla. A member of this family is called a soricine, or a red-toothed shrew. Soricinae is one of three subfamilies in Soricidae, along with the white-toothed shrews of Crocidurinae and the African shrews of Myosoricinae. They are found in Europe, Asia, North America, and northern South America, primarily in forests, shrublands, grasslands, and wetlands, and also in rocky areas and deserts. They range in size from the Eurasian pygmy shrew, at 3 cm (1 in) plus a 3 cm (1 in) tail, to various members of the Chimarrogale genus, at 14 cm (6 in) plus a 13 cm (5 in) tail. Soricines primarily eat insects and other invertebrates, as well as small vertebrates and plants. No soricines have population estimates, but 6 species are categorized as endangered species: the Bornean water shrew, Darién small-eared shrew, Enders's small-eared shrew, Guatemalan broad-clawed shrew, Sierra de Aroa shrew, and Pribilof Island shrew. Additionally, the Nelson's small-eared shrew, San Cristobal shrew, and Sclater's shrew are categorized as critically endangered.

Brown shrew
Southern water shrew (Neomys anomalus)

The 158 extant species of Soricinae are divided into 13 genera; 76 of them are in Sorex and 41 in Cryptotis, with the remaining 41 species divided between the other 11. A few extinct prehistoric Soricinae species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.[1]

Conventions

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IUCN Red List categories
Conservation status
 EX Extinct (0 species)
 EW Extinct in the wild (0 species)
 CR Critically Endangered (3 species)
 EN Endangered (6 species)
 VU Vulnerable (6 species)
 NT Near threatened (4 species)
 LC Least concern (103 species)
Other categories
 DD Data deficient (34 species)
 NE Not evaluated (1 species)

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the soricine's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.

Classification

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The shrew subfamily Soricinae consists of thirteen genera. The majority of the species are contained within Cryptotis, with 41 species, and Sorex, with 76 species. The remaining shrews are split between Chodsigoa with eight species; Chimarrogale with six; Anourosorex, Episoriculus, Neomys, and Notiosorex with four species each; Blarina and Blarinella with three species each, and Megasorex, Nectogale, and Soriculus each with a single species.

Subfamily Soricinae

  • Genus Anourosorex (Asian mole shrews): four species
  • Genus Blarina (short-tailed shrews): three species
  • Genus Blarinella (Asian short-tailed shrews): three species
  • Genus Chimarrogale (Asiatic water shrews): six species
  • Genus Chodsigoa (Asiatic shrews): eight species
  • Genus Cryptotis (small-eared shrews): 41 species
  • Genus Episoriculus (brown-toothed shrews): four species
  • Genus Megasorex (Mexican shrew): one species
  • Genus Nectogale (Elegant water shrew): one species
  • Genus Neomys (water shrews): four species
  • Genus Notiosorex (gray shrews): four species
  • Genus Sorex (long-tailed shrews): 76 species
  • Genus Soriculus (Himalayan shrew): one species
Soricinae[2][3][4]  

Soricines

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The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis, as supported by both the IUCN and the American Society of Mammalogists.[5]

Genus AnourosorexA. Milne-Edwards, 1872 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Assam mole shrew


A. assamensis
Anderson, 1875
Northeastern India
 
Size: 8–12 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[6]

Habitat: Forest and rocky areas[7]

Diet: Insects and earthworms[6]
 LC 


Unknown  [7]

Chinese mole shrew


A. squamipes
H. Milne-Edwards, 1872
Eastern Asia
 
Size: 8–11 cm (3–4 in), plus tail[8]

Habitat: Forest[9]

Diet: Insects and worms, as well as plants[8]
 LC 


Unknown  [9]

Giant mole shrew


A. schmidi
Petter, 1963
Northeastern India and Bhutan
 
Size: 8–12 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[6]

Habitat: Forest[10]

Diet: Insects and earthworms[6]
 DD 


Unknown  [10]

Taiwanese mole shrew

 

A. yamashinai
Kuroda, 1935
Taiwan
 
Size: 8–12 cm (3–5 in) long, plus 0.5–2 cm (0.2–0.8 in) tail[6]

Habitat: Forest and grassland[11]

Diet: Insects and earthworms[6]
 LC 


Unknown  [11]

Genus BlarinaGray, 1838 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Elliot's short-tailed shrew

 

B. hylophaga
Elliot, 1899
Central United States
 
Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[12]

Habitat: Grassland, shrubland, and forest[13]

Diet: Insects, arthropods, and earthworms, as well as small vertebrates and plants[12]
 LC 


Unknown  [13]

Everglades short-tailed shrew


B. peninsulae
Merriam, 1895
Florida Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[14]

Habitat: Forest, shrubland, grassland, and inland wetlands[14]

Diet: Invertebrates, small vertebrates, and plants[14]
 NE 


Unknown  

Northern short-tailed shrew

 

B. brevicauda
(Say, 1823)
Eastern North America
 
Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[15]

Habitat: Inland wetlands, shrubland, and forest[16]

Diet: Invertebrates, small vertebrates, and plants[15]
 LC 


Unknown  [16]

Southern short-tailed shrew

 

B. carolinensis
(Bachman, 1837)

Two subspecies
  • B. c. carolinensis
  • B. c. minima
Southeastern United States
 
Size: 7–11 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[17]

Habitat: Forest, shrubland, grassland, and inland wetlands[18]

Diet: Invertebrates, as well as plants[17]
 LC 


Unknown  [18]

Genus BlarinellaThomas, 1911 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Asiatic short-tailed shrew


B. quadraticauda
H. Milne-Edwards, 1872
Central China
 
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[19]

Habitat: Forest and shrubland[20]

Diet: Invertebrates[19]
 NT 


Unknown  [20]

Burmese short-tailed shrew


B. wardi
Thomas, 1915
Southern China and Myanmar
 
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[19]

Habitat: Forest[21]

Diet: Invertebrates[19]
 LC 


Unknown  [21]

Indochinese short-tailed shrew


B. griselda
Thomas, 1912
Central and southern China and northern Vietnam
 
Size: 6–9 cm (2–4 in) long, plus 3–6 cm (1–2 in) tail[19]

Habitat: Forest[22]

Diet: Invertebrates[19]
 LC 


Unknown  [22]

Genus ChimarrogaleAnderson, 1877 – six species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bornean water shrew


C. phaeura
Thomas, 1898
Northern Borneo in Malaysia
 
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest and inland wetlands[23]

Diet: Invertebrates[23]
 EN 


Unknown  [23]

Chinese water shrew


C. styani
De Winton, 1899
Central China and Myanmar
 
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest, shrubland, and inland wetlands[24]

Diet: Aquatic insects, crustaceans, and small fish[6]
 LC 


Unknown  [24]

Himalayan water shrew


C. himalayica
(Gray, 1842)
Eastern Asia
 
Size: 11–14 cm (4–6 in) long, plus 7–9 cm (3–4 in) tail[25]

Habitat: Forest and inland wetlands[26]

Diet: Insects and spiders[25]
 LC 


Unknown  [26]

Japanese water shrew

 

C. platycephalus
Temminck, 1842
Japan
 
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Inland wetlands[27]

Diet: Aquatic insects, crustaceans, and small fish[6]
 LC 


Unknown  [27]

Malayan water shrew


C. hantu
Harrison, 1958
Malaysia
 
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest and inland wetlands[28]

Diet: Aquatic insects, crustaceans, and small fish[6]
 NT 


Unknown  [28]

Sumatran water shrew


C. sumatrana
(Thomas, 1921)
Southern Sumatra in Indonesia
 
Size: 8–14 cm (3–6 in) long, plus 6–13 cm (2–5 in) tail[6]

Habitat: Forest and inland wetlands[29]

Diet: Aquatic insects, crustaceans, and small fish[29]
 DD 


Unknown  [29]

Genus ChodsigoaKastchenko, 1907 – eight species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
De Winton's shrew


C. hypsibia
De Winton, 1899
Central China
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest and shrubland[31]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [31]

Lamulate shrew


C. lamula
(Thomas, 1912)
Central China
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[32]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [32]

Lesser Taiwanese shrew


C. sodalis
(Thomas, 1913)
Taiwan
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[33]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown  [33]

Lowe's shrew


C. parca
(Allen, 1923)

Three subspecies
  • C. p. furva
  • C. p. lowei
  • C. p. parca
Southern Asia
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[34]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [34]

Pygmy brown-toothed shrew


C. parva
Allen, 1923
Southern China
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Unknown[35]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown  [35]

Salenski's shrew


C. salenskii
(Kastchenko, 1907)
Central China
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Unknown[36]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown  [36]

Smith's shrew


C. smithii
(Thomas, 1911)
Central China
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[37]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 NT 


Unknown  [37]

Van Sung's shrew


C. caovansunga
(Lunde, Musser, & Son, 2003)
Northern Vietnam
 
Size: 4–10 cm (2–4 in) long, plus 3–12 cm (1–5 in) tail[30]

Habitat: Forest[38]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 DD 


Unknown  [38]

Genus CryptotisPomel, 1848 – 41 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Big Mexican small-eared shrew


C. magna
(Merriam, 1895)
Southern Mexico
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[40]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 VU 


Unknown  [40]

Blackish small-eared shrew


C. nigrescens
(J. A. Allen, 1895)
Central America
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[41]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [41]

Blind small-eared shrew


C. niausa
Moreno Cárdenas & Albuja, 2014
Northern Ecuador Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest, grassland, and rocky areas[42]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [42]

Celaque broad-clawed shrew


C. celaque
Woodman, 2015
Western Honduras Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[43]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [43]

Central American least shrew

 

C. orophila
(Allen, 1895)
Central America
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest, shrubland, and inland wetlands[44]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [44]

Central Mexican broad-clawed shrew


C. alticola
(Merriam, 1895)
Central Mexico
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[45]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [45]

Colombian small-eared shrew


C. colombiana
Woodman & Timm, 1993
Central Colombia
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[46]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [46]

Darién small-eared shrew


C. merus
Goldman, 1912
Border of Colombia and Panama
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[47]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown  [47]

Dinira small-eared shrew


C. dinirensis
Quiroga-Carmona & DoNascimiento, 2016
Northern Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[48]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [48]

Eastern Cordillera small-footed shrew


C. brachyonyx
Woodman, 2003
Central Colombia
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Unknown[49]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [49]

Ecuadorian small-eared shrew


C. equatoris
Thomas, 1912
Ecuador
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and unknown[50]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [50]

Enders's small-eared shrew


C. endersi
Setzer, 1950
Panama
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[51]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown  [51]

Goldman's broad-clawed shrew


C. goldmani
(Merriam, 1895)
Southern Mexico
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[52]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [52]

Goodwin's broad-clawed shrew


C. goodwini
Jackson, 1933
Southern Mexico and Central America
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[53]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [53]

Grizzled Mexican small-eared shrew


C. obscura
(Merriam, 1895)
Eastern Mexico Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[54]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [54]

Guatemalan broad-clawed shrew


C. griseoventris
Jackson, 1933
Southern Mexico and Guatemala
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[55]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown  [55]

Honduran small-eared shrew


C. hondurensis
Woodman & Timm, 1992
Honduras
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[56]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [56]

Mam broad-clawed shrew


C. mam
Woodman, 2010
Guatemala Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[57]

Diet: Earthworms, insects, and plants[57]
 DD 


Unknown  [57]

Medellín small-eared shrew

 

C. medellinia
Thomas, 1921
Colombia
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[58]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [58]

Merida small-eared shrew


C. meridensis
Thomas, 1898
Venezuela
 
Size: 7–11 cm (3–4 in) long, plus 2–5 cm (1–2 in) tail[59]

Habitat: Forest and shrubland[60]

Diet: Insects and other invertebrates, as well as small vertebrates and eggs[59]
 VU 


Unknown  [60]

Merriam's small-eared shrew


C. merriami
Choate, 1970
Southern Mexico and Central America
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[61]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [61]

Mexican small-eared shrew


C. mexicana
(Coues, 1870)
Southern Mexico
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[62]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [62]

Muscular broad-clawed shrew


C. lacertosus
Woodman, 2010
Guatemala Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[63]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [63]

Nelson's small-eared shrew


C. nelsoni
Merriam, 1895
Eastern Mexico
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[64]

Diet: Insects[64]
 CR 


Unknown  [64]

North American least shrew

 

C. parva
(Say, 1823)
Eastern North America
 
Size: 7–10 cm (3–4 in) long, plus 1–3 cm (0.4–1.2 in) tail[65]

Habitat: Forest, shrubland, grassland, inland wetlands, and rocky areas[66]

Diet: Insects, as well as other invertebrates[65]
 LC 


Unknown  [66]

Oaxacan broad-clawed shrew


C. peregrina
(Merriam, 1895)
Southern Mexico Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[67]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [67]

Omoa broad-clawed shrew


C. mccarthyi
Woodman, 2015
Honduras Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[68]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [68]

Perija small-eared shrew


C. perijensis
Quiroga-Carmona & Woodman, 2015
Border of Colombia and Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[69]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [69]

Peruvian small-eared shrew


C. peruviensis
Vivar, Pacheco, & Valqui, 1997
Northern Peru
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[70]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [70]

Phillips's small-eared shrew


C. phillipsii
(Schaldach, 1966)
Southern Mexico Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[71]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 VU 


Unknown  [71]

Santa Barbara broad-clawed shrew


C. cavatorculus
Woodman, 2015
Honduras Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[72]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [72]

Scaly-footed small-eared shrew


C. squamipes
(Allen, 1916)
Colombia and Ecuador
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[73]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [73]

Sierra de Aroa shrew


C. aroensis
Quiroga-Carmona & Molinari, 2012
Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[74]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 EN 


Unknown  [74]

Talamancan small-eared shrew


C. gracilis
Miller, 1911
Costa Rica and Panama
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[75]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [75]

Tamá small-eared shrew


C. tamensis
Woodman, 2002
Border of Colombia and Venezuela
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[76]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [76]

Thomas's small-eared shrew


C. thomasi
(Merriam, 1897)
Colombia
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest and grassland[77]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [77]

Tropical small-eared shrew


C. tropicalis
(Merriam, 1895)
Southern Mexico and Central America
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest, grassland, and inland wetlands[78]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [78]

Venezuelan small-eared shrew


C. venezuelensis
Quiroga-Carmona, 2013
Northern Venezuela Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[79]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [79]

Wandering small-eared shrew


C. montivaga
(Anthony, 1921)
Ecuador
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[80]

Diet: Insects and other invertebrates[80]
 LC 


Unknown  [80]

Yalijux shrew


C. oreoryctes
Woodman, 2011
Guatemala Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[81]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 DD 


Unknown  [81]

Yucatan small-eared shrew


C. mayensis
(Merriam, 1901)
Southern Mexico and Central America
 
Size: 5–10 cm (2–4 in) long, plus 1–5 cm (0.4–2.0 in) tail[39]

Habitat: Forest[82]

Diet: Invertebrates, small lizards and frogs, and carrion[39]
 LC 


Unknown  [82]

Genus EpisoriculusEllerman & Morrison-Scott, 1966 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Hodgson's brown-toothed shrew


E. caudatus
(Horsfield, 1851)
Southern Asia
 
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest, shrubland, and grassland[83]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [83]

Long-tailed brown-toothed shrew


E. leucops
(Horsfield, 1855)
Southern Asia
 
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest[84]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [84]

Long-tailed mountain shrew


E. macrurus
Blanford, 1888
Southern Asia
 
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest[85]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [85]

Taiwanese brown-toothed shrew


E. fumidus
(Thomas, 1913)
Taiwan
 
Size: 4–7 cm (2–3 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest and shrubland[86]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [86]

Genus MegasorexHibbard, 1950 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Mexican shrew


M. gigas
(Merriam, 1897)
Southern Mexico
 
Size: 8–9 cm (3–4 in) long, plus 3–5 cm (1–2 in) tail[87]

Habitat: Forest and shrubland[88]

Diet: Invertebrates[87]
 LC 


Unknown  [88]

Genus NectogaleA. Milne-Edwards, 1870 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Elegant water shrew


N. elegans
H. Milne-Edwards, 1870
Central China and southern Asia
 
Size: 9–13 cm (4–5 in) long, plus 8–11 cm (3–4 in) tail[89]

Habitat: Forest and inland wetlands[90]

Diet: Insects, crustaceans, and small fish[89]
 LC 


Unknown  [90]

Genus NeomysKaup, 1829 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Eurasian water shrew

 

N. fodiens
(Pennant, 1771)
Europe and Asia
 
Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Forest, grassland, inland wetlands, intertidal marine, and coastal marine[91]

Diet: Aquatic invertebrates, fish, amphibians, and frogs[92]
 LC 


Unknown  [91]

Iberian water shrew


N. anomalus
A. Cabrera, 1907
Western Europe Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Inland wetlands[93]

Diet: Invertebrates and small vertebrates such as fish and frogs[87]
 LC 


Unknown  [93]

Mediterranean water shrew

 

N. milleri
Mottaz, 1907
Europe and western Asia
 
Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Inland wetlands[94]

Diet: Invertebrates and small vertebrates such as fish and frogs[87]
 LC 


Unknown  [94]

Transcaucasian water shrew


N. teres
Miller, 1908
Western Asia
 
Size: 6–10 cm (2–4 in) long, plus 4–8 cm (2–3 in) tail[87]

Habitat: Forest and inland wetlands[95]

Diet: Invertebrates, molluscs, fish, frogs, and small rodents[95]
 LC 


Unknown  [95]

Genus NotiosorexCoues, 1877 – four species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Cockrum's gray shrew

 

N. cockrumi
Baker, O'Neill, & McAliley, 2003
Northern Mexico and southern United States
 
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Shrubland and desert[97]

Diet: Insects and other invertebrates, as well as carrion[87]
 LC 


Unknown  [97]

Crawford's gray shrew

 

N. crawfordi
(Coues, 1877)
Southern North America
 
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Forest, shrubland, grassland, and desert[98]

Diet: Worms, spiders, and insects, as well as small mammals, lizards, and birds[99]
 LC 


Unknown  [98]

Large-eared gray shrew


N. evotis
(Coues, 1877)
Western Mexico
 
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Shrubland and grassland[100]

Diet: Insects and other invertebrates, as well as carrion[87]
 LC 


Unknown  [100]

Villa's gray shrew


N. villai
Carraway & Timm, 2000
Eastern Mexico
 
Size: 4–7 cm (2–3 in) long, plus 2–4 cm (1–2 in) tail[96]

Habitat: Forest[101]

Diet: Insects and other invertebrates, as well as carrion[87]
 VU 


Unknown  [101]

Genus SorexLinnaeus, 1758 – 76 species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Alaska tiny shrew


S. yukonicus
Dokuchaev, 1997
Alaska
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands[103]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [103]

Alpine shrew


S. alpinus
Schinz, 1837
Europe
 
Size: 6–8 cm (2–3 in) long, plus 5–8 cm (2–3 in) tail[105]

Habitat: Forest, grassland, and rocky areas[106]

Diet: Insects, arachnids, snails, and earthworms[105]
 NT 


Unknown  [106]

American pygmy shrew


S. hoyi
Baird, 1857
Northern North America
 
Size: 5–6 cm (2 in) long, plus 2–4 cm (1–2 in) tail[107]

Habitat: Forest, shrubland, grassland, and inland wetlands[108]

Diet: Insects and other invertebrates[107]
 LC 


Unknown  [108]

American water shrew

 

S. palustris
Richardson, 1828
North America
 
Size: 7–9 cm (3–4 in) long, plus 5–9 cm (2–4 in) tail[109]

Habitat: Forest and inland wetlands[110]

Diet: Aquatic insects and fish, as well as other invertebrates and plants[109]
 LC 


Unknown  [110]

Apennine shrew


S. samniticus
Altobello, 1926
Italy
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland and forest[111]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [111]

Arctic shrew


S. arcticus
Kerr, 1792
Northern North America
 
Size: 6–8 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[112]

Habitat: Forest and inland wetlands[113]

Diet: Insects and other invertebrates[112]
 LC 


Unknown  [113]

Arizona shrew


S. arizonae
Diersing & Hoffmeister, 1977
Northern Mexico and southern United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[114]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [114]

Azumi shrew


S. hosonoi
Imaizumi, 1954
Japan
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and grassland[115]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [115]

Baird's shrew

 

S. bairdi
Merriam, 1895
Northwestern United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[116]

Diet: Invertebrates[116]
 LC 


Unknown  [116]

Barren ground shrew


S. ugyunak
Anderson & Rand, 1945
Northern North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland[117]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [117]

Buchara shrew

 

S. buchariensis
Ognew, 1921
Central Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[118]

Diet: Insects[118]
 DD 


Unknown  [118]

Carmen Mountain shrew


S. milleri
Jackson, 1947
Northeastern Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[119]

Diet: Insects[119]
 VU 


Unknown  [119]

Caucasian pygmy shrew

 

S. volnuchini
Ognew, 1922
Western Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[120]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [120]

Caucasian shrew


S. satunini
Ognew, 1922
Western Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and rocky areas[121]

Diet: Insects[121]
 LC 


Unknown  [121]

Chestnut-bellied shrew


S. ventralis
Merriam, 1895
Southern Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[122]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [122]

Chinese highland shrew


S. excelsus
Allen, 1923
Southern China
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and grassland[123]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [123]

Chinese shrew


S. sinalis
Thomas, 1912
Central China
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Rocky areas[124]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [124]

Cinereus shrew

 

S. cinereus
Kerr, 1792
Northern North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, and inland wetlands[125]

Diet: A variety of invertebrates, as well as seeds and fungi[126]
 LC 


Unknown  [125]

Common shrew

 

S. araneus
Linnaeus, 1758
Europe and Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, inland wetlands, and coastal marine[127]

Diet: Insects and other invertebrates[128]
 LC 


Unknown  [127]

Crowned shrew

 

S. coronatus
Millet, 1828
Western Europe
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and inland wetlands[129]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [129]

Dwarf shrew


S. nanus
Merriam, 1895
Central United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Rocky areas, inland wetlands, grassland, and forest[130]

Diet: Insects and other invertebrates, as well as plants and small vertebrates[130]
 LC 


Unknown  [130]

Eurasian least shrew

 

S. minutissimus
Zimmermann, 1780
Norther Europe and northern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[131]

Diet: Insects and other invertebrates[131]
 LC 


Unknown  [131]

Eurasian pygmy shrew

 

S. minutus
Linnaeus, 1766
Europe and Asia
 
Size: 3–6 cm (1–2 in) long, plus 3–5 cm (1–2 in) tail[132]

Habitat: Forest, shrubland, grassland, inland wetlands, and coastal marine[133]

Diet: Insects and other invertebrates[132]
 LC 


Unknown  [133]

Flat-skulled shrew


S. roboratus
Hollister, 1913
Northern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[134]

Diet: Insects[134]
 LC 


Unknown  [134]

Fog shrew


S. sonomae
Jackson, 1921

Two subspecies
  • S. s. sonomae
  • S. s. tenelliodus
Western North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[135]

Diet: Insects and other invertebrates[135]
 LC 


Unknown  [135]

Gansu shrew


S. cansulus
Thomas, 1912
Central China
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Unknown[136]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [136]

Glacier Bay water shrew


S. alaskanus
Merriam, 1900
Western Canada
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands[137]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [137]

Greater stripe-backed shrew


S. cylindricauda
H. Milne-Edwards, 1872
Central China
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[138]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [138]

Iberian shrew

 

S. granarius
Miller, 1910
Portugal and Spain
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and rocky areas[139]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [139]

Inyo shrew

 

S. tenellus
Merriam, 1895
Western United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, inland wetlands, and rocky areas[140]

Diet: Insects and other invertebrates[140]
 LC 


Unknown  [140]

Ixtlan shrew


S. ixtlanensis
Carraway, 2007
Southern Mexico Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[141]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [141]

Jalisco shrew


S. mediopua
Carraway, 2007
Southern Mexico Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[142]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [142]

Kamchatka shrew


S. camtschatica
Yudin, 1972
Eastern Russia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland[143]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [143]

Kashmir pygmy shrew


S. planiceps
Miller, 1911
Northern India and Pakistan
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and rocky areas[144]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [144]

Kozlov's shrew


S. kozlovi
Stroganov, 1952
South-central China
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Unknown[145]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [145]

Large-toothed shrew


S. macrodon
Merriam, 1895
Southern Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[146]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 VU 


Unknown  [146]

Laxmann's shrew

 

S. caecutiens
Laxmann, 1788
Northern Europe and northern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and inland wetlands[147]

Diet: Insects and other invertebrates[147]
 LC 


Unknown  [147]

Lesser striped shrew


S. bedfordiae
Thomas, 1911
Central China and southern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[148]

Diet: Insects[148]
 LC 


Unknown  [148]

Long-clawed shrew

 

S. unguiculatus
Dobson, 1890
Eastern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[149]

Diet: Insects and other invertebrates[149]
 LC 


Unknown  [149]

Long-tailed shrew

 

S. dispar
Batchelder, 1911
Eastern North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[150]

Habitat: Forest, inland wetlands, and rocky areas[151]

Diet: Small invertebrates and plants[150]
 LC 


Unknown  [151]

Maritime shrew


S. maritimensis
Smith, 1939
Eastern Canada
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, inland wetlands, and unknown[152]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [152]

Marsh shrew

 

S. bendirii
Merriam, 1884
Western North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[153]

Diet: Insects[153]
 LC 


Unknown  [153]

Merriam's shrew

 

S. merriami
Dobson, 1890
Western United States
 
Size: 5–7 cm (2–3 in) long, plus 3–5 cm (1–2 in) tail[154]

Habitat: Forest, shrubland, and grassland[155]

Diet: Insects and other invertebrates, as well as small vertebrates[154]
 LC 


Unknown  [155]

Mexican long-tailed shrew


S. oreopolus
Merriam, 1892
Southern Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[156]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [156]

Montane shrew

 

S. monticolus
Merriam, 1890
Western North America
 
Size: 6–8 cm (2–3 in) long, plus 4–7 cm (2–3 in) tail[157]

Habitat: Forest, grassland, and inland wetlands[158]

Diet: Insects and other invertebrates, as well as salamanders, seeds, lichen, and fungi[157]
 LC 


Unknown  [158]

Mount Lyell shrew


S. lyelli
Merriam, 1902
Western United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland, grassland, and inland wetlands[159]

Diet: Insects[159]
 LC 


Unknown  [159]

New Mexico shrew


S. neomexicanus
Bailey, 1913
Southern United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[160]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [160]

Orizaba long-tailed shrew


S. orizabae
Merriam, 1895
Central Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[161]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [161]

Ornate shrew

 

S. ornatus
Merriam, 1895

Eight subspecies
  • S. o. juncensis (Tule shrew)
  • S. o. lagunae
  • S. o. ornatus
  • S. o. relictus
  • S. o. salarius
  • S. o. salicornicus
  • S. o. willetti
  • S. o. sinuosus (Suisun shrew)
Western North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, and inland wetlands[162]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [162]

Pacific shrew


S. pacificus
Coues, 1877

Two subspecies
  • S. p. cascadensis
  • S. p. pacificus
Western United States
 
Size: 8–9 cm (3–4 in) long, plus 5–8 cm (2–3 in) tail[163]

Habitat: Forest[164]

Diet: Insects, other invertebrates, and amphibians, as well as plants and fungi[163]
 LC 


Unknown  [164]

Paramushir shrew


S. leucogaster
Kuroda, 1933
Eastern Russia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Shrubland[165]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [165]

Portenko's shrew


S. portenkoi
Stroganov, 1956
Eastern Russia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland[166]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [166]

Prairie shrew


S. haydeni
Baird, 1857
Central North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, and inland wetlands[167]

Diet: Insects and other invertebrates, as well as small vertebrates and plants[167]
 LC 


Unknown  [167]

Preble's shrew


S. preblei
Jackson, 1922
Northwestern United States
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, grassland, inland wetlands, and desert[168]

Diet: Insects[168]
 LC 


Unknown  [168]

Pribilof Island shrew


S. pribilofensis
Merriam, 1895
Pribilof Islands in Alaska
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland and coastal marine[169]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 EN 


Unknown  [169]

Radde's shrew


S. raddei
Satunin, 1895
Western Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and rocky areas[170]

Diet: Insects[170]
 LC 


Unknown  [170]

Saint Lawrence Island shrew


S. jacksoni
Hall & Gilmore, 1932
St. Lawrence Island in Alaska
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Grassland, inland wetlands, and rocky areas[171]

Diet: Insects[171]
 LC 


Unknown  [171]

San Cristobal shrew


S. stizodon
Merriam, 1895
Southeastern Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[172]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 CR 


Unknown  [172]

Saussure's shrew


S. saussurei
Merriam, 1892
Southern Mexico and Guatemala
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[173]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [173]

Sclater's shrew


S. sclateri
Merriam, 1897
Southern Mexico and Guatemala
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[174]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 CR 


Unknown  [174]

Shinto shrew


S. shinto
Thomas, 1905

Three subspecies
  • S. s. sadonis (Sado shrew)
  • S. s. shikokensis
  • S. s. shinto
Japan
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[175]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [175]

Siberian large-toothed shrew


S. daphaenodon
Thomas, 1907
Northern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands and forest[176]

Diet: Insects and other invertebrates[176]
 LC 


Unknown  [176]

Slender shrew


S. gracillimus
Thomas, 1907

Four subspecies
  • S. g. gracillimus
  • S. g. granti
  • S. g. minor
  • S. g. natalae
Eastern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and grassland[177]

Diet: Invertebrates[177]
 LC 


Unknown  [177]

Smoky shrew

 

S. fumeus
Miller, 1895
Eastern North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[178]

Diet: Invertebrates, as well as salamanders and fungi[179]
 LC 


Unknown  [178]

Southeastern shrew

 

S. longirostris
Bachman, 1837

Three subspecies
  • S. l. eonis
  • S. l. fisheri (Dismal Swamp southeastern shrew)
  • S. l. longirostris
Eastern United States
 
Size: 4–6 cm (2–2 in) long, plus 2–4 cm (1–2 in) tail[180]

Habitat: Inland wetlands, grassland, shrubland, and forest[181]

Diet: Insects and other invertebrates as well as seeds[180]
 LC 


Unknown  [181]

Taiga shrew


S. isodon
Turov, 1924
Northern Europe and northern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[182]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [182]

Tibetan shrew


S. thibetanus
Kaschtschenko, 1905
Central China
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and grassland[183]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [183]

Tien Shan shrew


S. asper
Thomas, 1914
Central Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and shrubland[184]

Diet: Insects and other invertebrates[184]
 LC 


Unknown  [184]

Trowbridge's shrew

 

S. trowbridgii
Baird, 1857
Western North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest, shrubland, and inland wetlands[185]

Diet: Insects and other invertebrates[185]
 LC 


Unknown  [185]

Tundra shrew

 

S. tundrensis
Merriam, 1900
Asia and western North America
 
Size: 8–12 cm (3–5 in) long, plus 2–4 cm (1–2 in) tail[186]

Habitat: Forest, grassland, and inland wetlands[187]

Diet: Insects, other invertebrates, and flowers[186]
 LC 


Unknown  [187]

Ussuri shrew


S. mirabilis
Ognew, 1937
Eastern Asia
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[188]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 DD 


Unknown  [188]

Vagrant shrew

 

S. vagrans
Baird, 1857
Western North America
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest and inland wetlands[189]

Diet: Insects and other invertebrates[189]
 LC 


Unknown  [189]

Valais shrew

 

S. antinorii
Bonaparte, 1840
Southern Europe
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Inland wetlands[190]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [190]

Veracruz shrew


S. veraecrucis
Jackson, 1925
Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[191]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [191]

Verapaz shrew


S. veraepacis
Alston, 1877

Three subspecies
  • S. v. chiapensis
  • S. v. mutabilis
  • S. v. veraepacis
Southern Mexico and Guatemala
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[192]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [192]

Zacatecas shrew


S. emarginatus
Jackson, 1925
Central Mexico
 
Size: 4–10 cm (2–4 in) long, plus 2–9 cm (1–4 in) tail[102]

Habitat: Forest[193]

Diet: Insects, worms, and other invertebrates, as well as small vertebrates, carrion, and plants[104]
 LC 


Unknown  [193]

Genus SoriculusBlyth, 1854 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Himalayan shrew

 

S. nigrescens
(Gray, 1842)
Himalayas in Asia
 
Size: 8–10 cm (3–4 in) long, plus 3–7 cm (1–3 in) tail[30]

Habitat: Forest, shrubland, and rocky areas[194]

Diet: Earthworms, insects, and other invertebrates, as well as small mammals[30]
 LC 


Unknown  [194]

References

edit
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