Euhadra

Euhadra
	Euhadra quaesita
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Order:	Stylommatophora
Family:	Camaenidae
Subfamily:	Bradybaeninae
Tribe:	Euhadrini
Genus:	Euhadra; Pilsbry, 1890
Synonyms
	Eulota (Euhadra) Pilsbry, 1890; Helix (Euhadra) Pilsbry, 1890;

Euhadra is a genus of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily Bradybaeninae of the family Camaenidae.^[2]

(This snail genus was previously placed in the family Eulotidae).

A few of the species in this genus are unusual in that specimens in those species always have left-handed "sinistral" coiling in their shells, as shown in the specimen on the right. The rest of the species in the genus are right-handed or "dextral" in the shell coiling, as is usually the case in the great majority of gastropods.

Distribution

These snails are almost endemic to Japan, but one of them are distributed in Korea.

Species

Most species in this genus have a dextral shell and 5 species are sinistral.^[3]

Dextral species

Dextral species in the genus Euhadra include:

Euhadra amaliae (Kobelt, 1875)
Euhadra awaensis (Pilsbry, 1902)
Euhadra brandtii (Kobelt, 1875)
Euhadra callizona (Crosse, 1871)
Euhadra caspari (Möllendorff, 1884)
Euhadra cecillei (Philippi, 1849)
Euhadra congenita (E. A. Smith, 1878)
Euhadra cyclolabris Möllendorff, 1899
Euhadra dixoni (Pilsbry, 1900)
Euhadra eoa (Crosse, 1868)
Euhadra granulifera (Möllendorff, 1888)
Euhadra herklotsi (Martens, 1861)
Euhadra idzumonis (Pilsbry & Gulick, 1900)
Euhadra kunoensis Kuroda in Masuda & Habe, 1989
Euhadra latispira (Pilsbry & Hirase, 1909)
- Euhadra latispira yagurai Kuroda & Habe, 1949 [1] Archived 8 January 2009 at the Wayback Machine
Euhadra luhuana (G. B. Sowerby I, 1839)
Euhadra moreletiana (Heude, 1882)
Euhadra nachicola Kuroda, 1929
Euhadra nesiotica (Pilsbry, 1902)
† Euhadra pachya (Pilsbry, 1902)
Euhadra peliomphala (L. Pfeiffer, 1850)
Euhadra sadoensis (Pilsbry & Y. Hirase, 1903)
Euhadra sandai (Kobelt, 1879)
Euhadra schmackeri (Möllendorff, 1888)
Euhadra senckenbergiana (Kobelt, 1875)
- Euhadra senckenbergiana aomoriensis (Gulick & Pilsbry, 1900)
- Euhadra senckenbergiana ibukicola
- Euhadra senckenbergiana notoensis
- Euhadra senckenbergiana minoensis
- Euhadra senckenbergiana senckenbergiana
Euhadra sigeonis Kuroda, 1944
Euhadra subnimbosa (Kobelt, 1894)
† Euhadra takarajimana M. Azuma & Y. Azuma, 1985
Euhadra tokarainsula Minato & Habe, 1982
Euhadra yakushimana (Pilsbry & Hirase, 1903)

Sinistral species

Euhadra murayamai

Sinistral species in the genus Euhadra include:

Euhadra decorata (Pilsbry & Hirase, 1903)
Euhadra grata (Gude, 1900)
- Euhadra grata echigoensis Murayama, Takizawa & Habe, 1991
Euhadra murayamai Habe, 1976
Euhadra quaesita (Deshayes, 1850), Sought-after False Hadra
Euhadra scaevola (Martens, 1877)

Species brought into synonymy

Euhadra amphidroma Möllendorff, 1899: synonym of Trichocathaica amphidroma (Möllendorff, 1899) (original combination)
Euhadra carphochroa Möllendorff, 1899: synonym of Bradybaena carphochroa (Möllendorff, 1899) (original combination)
Euhadra eris Möllendorff, 1899: synonym of Bradybaena eris (Möllendorff, 1899) (original combination)
Euhadra halpozona Möllendorff, 1899: synonym of Bradybaena halpozona (Möllendorff, 1899) (original combination)
Euhadra hickonis Kobelt, 1879: synonym of Euhadra congenita (E. A. Smith, 1878) (junior synonym)
Euhadra iwadensis Toba & Kuroda, 1936: synonym of Euhadra decorata decorata (Pilsbry & Hirase, 1903) (junior synonym)
Euhadra kanmuriensis Azuma & Tada, 1969: synonym of Euhadra sandai communis Pilsbry, 1928 (junior synonym)
Euhadra kanoi Kuroda, 1932: synonym of Nesiohelix kanoi (Kuroda, 1932) (original combination)
Euhadra koreana (L. Pfeiffer, 1850): synonym of Fruticicola koreana (L. Pfeiffer, 1850)
Euhadra micromphala Möllendorff, 1899: synonym of Bradybaena micromphala (Möllendorff, 1899) (original combination)
Euhadra pekanensis H. Rolle, 1911: synonym of Satsuma pekanensis (H. Rolle, 1911) (original combination)
Euhadra pseudocampylaea Möllendorff, 1899: synonym of Bradybaena pseudocampylaea (Möllendorff, 1899) (original combination)
Euhadra pseudopapuina Möllendorff, 1901: synonym of Aegista pseudopapuina (Möllendorff, 1901) (original combination)
Euhadra strauchiana Möllendorff, 1899: synonym of Bradybaena strauchiana (Möllendorff, 1899) (original combination)
Euhadra strictotaenia Möllendorff, 1899: synonym of Bradybaena strictotaenia (Möllendorff, 1899) (original combination)
Euhadra tenuitesta Möllendorff, 1899: synonym of Bradybaena tenuitesta (Möllendorff, 1899) (original combination)
Euhadra tobai S. Hirase, 1929: synonym of Euhadra decorata decorata (Pilsbry & Hirase, 1903) (junior synonym)
Euhadra tokarana Azuma, 1982: synonym of Euhadra tokarainsula tokarainsula Minato & Habe, 1982

References

^ Pilsbry H. A. (1890) In: Tryon & Pilsbry, Manual of Conchology (2)6: 94-95.
^ MolluscaBase eds. (2021). MolluscaBase. Euhadra Pilsbry, 1890. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=995784 on 2021-02-08
^ Davison A, Chiba S, Barton NH, Clarke B (2005). "Speciation and Gene Flow between Snails of Opposite Chirality". PLOS Biology. 3 (9, e282): e282. doi:10.1371/journal.pbio.0030282. PMC 1182688. PMID 16149849.