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Rudolf Albert Raff (November 10, 1941 – January 5, 2019) was an American biologist, and James H. Rudy Professor of Biology at Indiana University.[4] He was known for research in, and promotion of, evolutionary developmental biology. He was also director of the Indiana Molecular Biology Institute.[5][6]

Rudy Raff
Rudolf A. Raff, June 2011.jpg
Raff in 2011
Born(1941-11-10)November 10, 1941
DiedJanuary 5, 2019(2019-01-05) (aged 77)
Bloomington, Indiana, United States
Alma mater
Scientific career
FieldsEvolutionary developmental biology
InstitutionsIndiana University, National Naval Medical Center, Massachusetts Institute of Technology


Raff was born in Shawnigan, Quebec[7] in 1941 to a family of Jewish immigrants from Eastern Europe. He graduated from Pennsylvania State University with a B.S. in 1963, and from Duke University with a Ph.D. in 1967. He died in 2019 in Bloomington Hospital, Indiana, at the age of 77.[8]



  • with Thomas C. Kaufman, Illustrated by E.C. Raff, Embryos, Genes, and Evolution: The Developmental-Genetic Basis of Evolutionary Change, Macmillan 1983, ISBN 0-02-397500-8
  • The shape of life: genes, development, and the evolution of animal form, University of Chicago Press, 1996, ISBN 978-0-226-70266-7
  • William R. Jeffery, Rudolf A. Raff (eds), Time, space, and pattern in embryonic development, A.R. Liss, 1983, ISBN 978-0-8451-2201-3
  • Rudolf A. Raff, Once We All Had Gills, Growing Up Evolutionist in an Evolving World, Indiana University Press 2012, ISBN 978-0-253-00235-8


  1. ^ "Historic Fellows".
  2. ^ "Rudolf A. Raff | American Academy of Arts and Sciences".
  3. ^ "John Simon Guggenheim Foundation | Rudolf A. Raff".
  4. ^ "Archived copy". Archived from the original on 2012-03-27. Retrieved 2011-10-29.CS1 maint: Archived copy as title (link)
  5. ^ "Archived copy". Archived from the original on 2012-07-22. Retrieved 2011-10-29.CS1 maint: Archived copy as title (link)
  6. ^ Akst, Jef (Jan 9, 2019). "Leader of Evo-Devo Field, Rudy Raff, Dies". The Scientist.
  7. ^
  8. ^ "Rudolf "Rudy" Albert Raff". Dignity Memorial.
  9. ^
  10. ^ McPeek, Mark (2005). "2004 Sewall Wright Award: Rudolf A. Raff". The American Naturalist. 165 (1): i. doi:10.1086/427345.
  11. ^ Lewis, Ricki (May 13, 2002). "Rudolf Raff". The Scientist.
  12. ^ Mikhailov, AT; Gilbert, SF (2002). "From development to evolution: the re-establishment of the "Alexander Kowalevsky Medal"". The International Journal of Developmental Biology. 46 (5): 693–8. PMID 12216980.
  13. ^

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