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The Moraceae — often called the mulberry family or fig family — are a family of flowering plants comprising about 38 genera and over 1180 species.[2] Most are widespread in tropical and subtropical regions, less so in temperate climates. The only synapomorphy within Moraceae is presence of laticifers and milky sap in all parenchymatous tissues, but generally useful field characters include two carpels sometimes with one reduced, compound inconspicuous flowers, and compound fruits.[3] The family includes well-known plants such as the fig, banyan, breadfruit, mulberry, and Osage-orange. The 'flowers' of Moraceae are often pseudanthia (reduced inflorescences).

Moraceae
Temporal range: 80–0 Ma
Cretaceous - Recent
Castilla elastica - Köhler–s Medizinal-Pflanzen-174.jpg
Panama rubber tree (Castilla elastica)
Scientific classification e
Kingdom: Plantae
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Rosales
Family: Moraceae
Gaudich.[1]
Genera

See text

Contents

ClassificationEdit

Formerly included within the now defunct order Urticales, recent molecular studies have resulted in its placement within Rosales in a clade called the urticalean rosids that also includes Ulmaceae, Celtidaceae, Cannabaceae and Urticaceae. Cecropia, which has variously been placed in Moraceae, Urticaceae, or their own family, Cecropiaceae, is now included in Urticaceae.[4]

Dioecy (having individuals with separate sexes) appears to be the primitive state in Moraceae.[5] Monoecy has evolved independently at least four times within the family.

Tribes and generaEdit

Moraceae comprises the following:[5][6]

PhylogenyEdit

Modern molecular phylogenetics suggest the following relationships:[4][5][6][7]



Urticaceae (outgroup)


Moraceae

Artocarpeae


Batocarpus



Clarisia





Artocarpus




Parartocarpus



Prainea





Moreae

Sorocea




Bagassa





Milicia



Streblus





Morus



Trophis








Maclureae

Maclura



Dorstenieae

Fatoua





Broussonetia



Malaisia






Bleekrodea



Sloetia




Trilepisium





Utsetela



Dorstenia





Brosimum



Trymatococcus



Helianthostylis








Ficeae

Ficus


Castilleae
Antiaropsineae

Sparattosyce



Antiaropsis



Castillineae


Antiaris



Mesogyne





Naucleopsis




Perebea



Pseudolmedia




Maquira




Helicostylis




Poulsenia



Castilla














ReferencesEdit

  1. ^ Angiosperm Phylogeny Group (2009). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III" (PDF). Botanical Journal of the Linnean Society. 161 (2): 105–121. doi:10.1111/j.1095-8339.2009.00996.x. Retrieved 2013-07-06. 
  2. ^ Christenhusz, M. J. M.; Byng, J. W. (2016). "The number of known plants species in the world and its annual increase". Phytotaxa. Magnolia Press. 261 (3): 201–217. doi:10.11646/phytotaxa.261.3.1. 
  3. ^ Judd WS, Campbell CS, Kellogg EA, Stevens PF, Donoghue MJ. (2008). Plant Systematics: A Phylogenetic Approach. Sunderland, MA: Sinauer Associates, Inc. pp. 1–620. ISBN 978-0-878-93407-2. 
  4. ^ a b Sytsma KJ, Morawetz J, Pires C, Nepokroeff M, Conti E, Zjhra M, Hall JC, Chase MW. (2002). "Urticalean rosids: Circumscription, rosid ancestry, and phylogenetics based on rbcL, trnLF, and ndhF sequences" (PDF). American Journal of Botany. 89 (89): 1531–1546. doi:10.3732/ajb.89.9.1531. PMID 21665755. 
  5. ^ a b c Datwyler SL, Weiblen G. (2004). "On the origin of the fig: Phylogenetic relationships of Moraceae from ndhF sequences". American Journal of Botany. 91 (5): 767–777. doi:10.3732/ajb.91.5.767. PMID 21653431. 
  6. ^ a b Clement WL, Weiblen GD. (2009). "Morphological evolution in the mulberry family (Moraceae)". Systematic Botany. 34 (3): 530–552. doi:10.1600/036364409789271155. 
  7. ^ Zerega NJC, Clement WL, Datwyler SL, Weiblen GD. (2005). "Biogeography and divergence times in the mulberry family (Moraceae)". Molecular Phylogenetics and Evolution. 37 (2): 402–416. doi:10.1016/j.ympev.2005.07.004. PMID 16112884. 

External linksEdit