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  1. ^ Although the plains and mountain zebras have been placed in Hippotigris and Grévy's zebra in Dolichohippus, Vilstrup et al. (2013) finds that the plains zebra and Grévy's zebra are more closely related.[4] Groves and Bell (2004) place all three species in the subgenus Hippotigris.[5]


  1. ^ Hay, Oliver P. (1915), Contributions to the Knowledge of the Mammals of the Pleistocene of North America, 48, Proceedings of the United States National Museum, pp. 535–549, doi:10.5479/si.00963801.48-2086.515
  2. ^ Aouraghe, Hassan; et al. (1999). "Les équidés du Pléstocène supérior de la grotte Zouhrah à El Harhoura, Maroc". Quaternaire (in French). 10: 283–292.
  3. ^ Orlando, L.; et al. (2008). "Ancient DNA Clarifies the Evolutionary History of American Late Pleistocene Equids". Journal of Molecular Evolution. 66 (5): 533–538. Bibcode:2008JMolE..66..533O. doi:10.1007/s00239-008-9100-x. PMID 18398561.
  4. ^ Vilstrup, Julia T.; et al. (2013). "Mitochondrial Phylogenomics of Modern and Ancient Equids". PLOS ONE. 8 (2): e55950. Bibcode:2013PLoSO...855950V. doi:10.1371/journal.pone.0055950. PMC 3577844. PMID 23437078.
  5. ^ Groves, C. P.; Bell, C. H. (2004). "New investigations on the taxonomy of the zebras genus Equus, subgenus Hippotigris". Mammalian Biology. 69 (3): 182–196. doi:10.1078/1616-5047-00133.
  6. ^ Eisenmann, Vera (March 2010). "Sussemionus, a new subgenus of Equus (Perissodactyla, Mammalia)". Comptes Rendus Biologies. 333 (3): 235–240. doi:10.1016/j.crvi.2009.12.013.
  7. ^ Eisenmann, Vera; Vasilijiev, Sergej (September 2011). "Unexpected finding of a new Equus species (Mammalia, Perissodactyla) belonging to a supposedly extinct subgenus in late Pleistocene deposits of Khakassia (Southwestern Siberia)". Geodiversitas. 33 (3): 519–530. doi:10.5252/g2011n3a5.
  8. ^ "Fossilworks: Equus fraternus". Retrieved 2016-04-23.