List of Odonata species of Australia

This is a list of species of damselflies and dragonflies recorded in Australia.

Alpine darner, Austroaeschna flavomaculata

Common names of species are linked, beside their scientific names.

The list is split into two groups: damselflies (suborder Zygoptera) and other dragonflies (infraorder Anisoptera). Those groups are organized in Families and then Genera and Species.

Zygoptera (damselflies)

Argiolestidae

genus: Archiargiolestes

• Midget flatwing, Archiargiolestes parvulus
• Tiny flatwing, Archiargiolestes pusillissimus
• Little flatwing, Archiargiolestes pusillus

genus: Austroargiolestes

• New England flatwing, Austroargiolestes alpinus
• Flame flatwing, Austroargiolestes amabilis
• Tropical flatwing, Austroargiolestes aureus
• Barrington flatwing, Austroargiolestes brookhousei
• Powdered flatwing, Austroargiolestes calcaris
• Milky flatwing, Austroargiolestes christine
• Golden flatwing, Austroargiolestes chrysoides
• Azure flatwing, Austroargiolestes elke
• Common flatwing, Austroargiolestes icteromelas
• Sydney flatwing, Austroargiolestes isabellae

genus: Griseargiolestes

• Coastal flatwing, Griseargiolestes albescens
• Turquoise flatwing, Griseargiolestes bucki
• Grey-chested flatwing, Griseargiolestes eboracus
• Springs flatwing, Griseargiolestes fontanus
• Grey flatwing, Griseargiolestes griseus
• Alpine flatwing, Griseargiolestes intermedius
• Metallic flatwing, Griseargiolestes metallicus

genus: Miniargiolestes

• Stream flatwing, Miniargiolestes minimus

genus: Podopteryx

• Treehole flatwing, Podopteryx selysi

Coenagrionidae

genus: Aciagrion

• Blue slim, Aciagrion fragilis

genus: Agriocnemis

• Silver wisp, Agriocnemis argentea
• Tropical wisp, Agriocnemis dobsoni
• Pilbara wisp, Agriocnemis kunjina
• Pygmy wisp, Agriocnemis pygmaea
• Red-rumped wisp, Agriocnemis rubricauda
• Agriocnemis thoracalis

genus: Archibasis

• Blue-banded longtail, Archibasis mimetes

genus: Argiocnemis

• Red-tipped shadefly, Argiocnemis rubescens

genus: Austroagrion

• South-western billabongfly, Austroagrion cyane
• Northern billabongfly, Austroagrion exclamationis
• Pilbara billabongfly, Austroagrion pindrina
• Eastern billabongfly, Austroagrion watsoni

genus: Austrocnemis

• Tiny longlegs, Austrocnemis maccullochi
• Kimberley longlegs, Austrocnemis obscura
• Splendid longlegs, Austrocnemis splendida

genus: Caliagrion

• Large riverdamsel, Caliagrion billinghursti

genus: Ceriagrion

• Redtail, Ceriagrion aeruginosum

genus: Coenagrion

• Swamp bluet, Coenagrion lyelli

genus: Ischnura

• Aurora bluetail, Ischnura aurora
• Common bluetail, Ischnura heterosticta
• Colourful bluetail, Ischnura pruinescens

genus: Pseudagrion

• Gold-fronted riverdamsel, Pseudagrion aureofrons
• Northern riverdamsel, Pseudagrion cingillum
• Flame-headed riverdamsel, Pseudagrion ignifer
• Dusky riverdamsel, Pseudagrion jedda
• Citrine-headed riverdamsel, Pseudagrion lucifer
• Blue riverdamsel, Pseudagrion microcephalum

genus: Teinobasis

• Red-breasted longtail, Teinobasis rufithorax

genus: Xanthagrion

• Red and blue damsel, Xanthagrion erythroneurum

Hemiphlebiidae

genus: Hemiphlebia

• Ancient greenling, Hemiphlebia mirabilis

Isostictidae

genus: Austrosticta

• Northern pondsitter, Austrosticta fieldi
• Eastern pondsitter, Austrosticta frater
• Kimberley pondsitter, Austrosticta soror

genus: Eurysticta

• Pilbara pin, Eurysticta coolawanyah
• Coomalie pin, Eurysticta coomalie
• Kimberley pin, Eurysticta kununurra
• Queensland pin, Eurysticta reevesi

genus: Labidiosticta

• Large wiretail, Labidiosticta vallisi

genus: Lithosticta

• Rock narrow-wing, Lithosticta macra

genus: Neosticta

• Southern pinfly, Neosticta canescens
• Tropical pinfly, Neosticta fraseri
• Forest pinfly, Neosticta silvarum

genus: Oristicta

• Slender wiretail, Oristicta filicicola
• Elegant wiretail, Oristicta rosendaleorum

genus: Rhadinosticta

• Northern wiretail, Rhadinosticta banksi
• Wiretail, Rhadinosticta handschini
• Powdered wiretail, Rhadinosticta simplex

Lestidae

genus: Austrolestes

• Western ringtail, Austrolestes aleison
• Slender ringtail, Austrolestes analis
• Blue ringtail, Austrolestes annulosus
• Inland ringtail, Austrolestes aridus
• Metallic ringtail, Austrolestes cingulatus
• Northern ringtail, Austrolestes insularis
• Iota ringtail, Austrolestes io
• Wandering ringtail, Austrolestes leda
• Dune ringtail, Austrolestes minjerriba
• Cup ringtail, Austrolestes psyche

genus: Indolestes

• Small reedling, Indolestes alleni
• Cave reedling, Indolestes obiri
• Slender reedling, Indolestes tenuissimus

genus: Lestes

• Dusky spreadwing, Lestes concinnus

Lestoideidae

genus: Diphlebia

• Sapphire rockmaster, Diphlebia coerulescens
• Tropical rockmaster, Diphlebia euphaeoides
• Giant rockmaster, Diphlebia hybridoides
• Whitewater rockmaster, Diphlebia lestoides
• Arrowhead rockmaster, Diphlebia nymphoides

genus: Lestoidea

• Large bluestreak, Lestoidea barbarae
• Short-tipped bluestreak, Lestoidea brevicauda
• Common bluestreak, Lestoidea conjuncta
• Mount Lewis bluestreak, Lestoidea lewisiana

Platycnemididae

genus: Nososticta

• Black-winged threadtail, Nososticta baroalba
• Green-blue threadtail, Nososticta coelestina
• Northern threadtail, Nososticta fraterna
• Spot-winged threadtail, Nososticta kalumburu
• Koolpinyah threadtail, Nososticta koolpinyah
• Citrine threadtail, Nososticta koongarra
• Malachite threadtail, Nososticta liveringa
• Striped threadtail, Nososticta mouldsi
• Pilbara threadtail, Nososticta pilbara
• Orange threadtail, Nososticta solida
• Fivespot threadtail, Nososticta solitaria
• Melville Island threadtail, Nososticta taracumbi

Synlestidae

genus: Chorismagrion

• Pretty relict, Chorismagrion risi

genus: Episynlestes

• Southern whitetip, Episynlestes albicauda
• Tropical whitetip, Episynlestes cristatus
• Intermediate whitetip, Episynlestes intermedius

genus: Synlestes

• Forest needle, Synlestes selysi
• Tropical needle, Synlestes tropicus
• Bronze needle, Synlestes weyersii

Anisoptera (dragonflies)

Aeshnidae

genus: Acanthaeschna

• Thylacine darner, Acanthaeschna victoria (Coastal southern Queensland and northern New South Wales)

genus: Adversaeschna

• Blue-spotted hawker, Adversaeschna brevistyla (Coastal areas from Cooktown in northern Queensland, south to Victoria, and the Pilbara in Western Australia)

genus: Agyrtacantha

• Trifid duskhawker, Agyrtacantha dirupta (Cape York Peninsula)

genus: Anaciaeschna

• Australasian duskhawker, Anaciaeschna jaspidea (Coastal Queensland and Northern Territory)

genus: Anax

• Kimberley emperor, Anax georgius (The Kimberley)
• Green emperor, Anax gibbosulus (Coastal northern Australia from the Kimberley to the Gold Coast)
• Lesser green emperor, Anax guttatus (Coastal northern Australia from the Kimberley to the Gold Coast)
• Australian emperor, Anax papuensis (All Australia and islands)

genus: Antipodophlebia

• Terrestrial evening darner, Antipodophlebia asthenes

genus: Austroaeschna

Lesser Tasmanian darner Austroaeschna hardyi

• Western darner, Austroaeschna anacantha (Coastal south-western Western Australia)
• Mountain darner, Austroaeschna atrata (Australian Alps)
• S-spot darner, Austroaeschna christine (Central coast of Queensland)
• Wallum darner, Austroaeschna cooloola Southern coast of Queensland
• Eungella darner, Austroaeschna eungella (Central coast of Queensland)
• Alpine darner, Austroaeschna flavomaculata (Australian Alps)
• Lesser Tasmanian darner, Austroaeschna hardyi (Tasmania)
• Whitewater darner, Austroaeschna inermis (Coastal south-eastern Australia)
• Grampians darner, Austroaeschna ingrid (Western Victoria)
• Carnarvon darner, Austroaeschna muelleri (Inland southern Queensland)
• Multi-spotted darner, Austroaeschna multipunctata (Southern coastal New South Wales to Victoria)
• Sydney mountain darner, Austroaeschna obscura (Mountains around Sydney)
• Swamp darner, Austroaeschna parvistigma (Coastal New South Wales, Victoria, southern South Australia and Tasmania)
• Inland darner, Austroaeschna pinheyi (Carnarvon Gorge, Queensland)
• Forest darner, Austroaeschna pulchra (Coastal southern Queensland, New South Wales and Victoria)
• Sigma darner, Austroaeschna sigma (Coastal southern Queensland and New South Wales)
• Tropical unicorn darner, Austroaeschna speciosa (Central coast of Queensland)
• Conehead darner, Austroaeschna subapicalis (Coastal New South Wales and Victoria)
• Tasmanian darner, Austroaeschna tasmanica (Tasmania)
• Unicorn darner, Austroaeschna unicornis (Southern Queensland to coastal New South Wales, Victoria, Tasmania and South Australia)

genus: Austrogynacantha

• Australian duskhawker, Austrogynacantha heterogena (Across Australia north of about Sydney)

genus: Austrophlebia

• Southern giant darner, Austrophlebia costalis
• Northern giant darner, Austrophlebia subcostalis

genus: Dendroaeschna

• Wide-faced darner, Dendroaeschna conspersa

genus: Dromaeschna

• Green-striped darner, Dromaeschna forcipata (Coast of Cape York Peninsula, Queensland)
• Ochre-tipped darner, Dromaeschna weiskei (Coast of Cape York Peninsula, Queensland)

genus: Gynacantha

• Lesser duskhawker, Gynacantha dobsoni (Northern Australia)
• Slender duskhawker, Gynacantha kirbyi (Northern Queensland)
• Paddle-tipped duskhawker, Gynacantha mocsaryi (Northern Queensland)
• Cave duskhawker, Gynacantha nourlangie (Inland and northern Australia)
• Grey duskhawker, Gynacantha rosenbergi (Northern Queensland)

genus: Notoaeschna

• Northern riffle darner, Notoaeschna geminata
• Southern riffle darner, Notoaeschna sagittata

genus: Spinaeschna

• Southern cascade darner, Spinaeschna tripunctata
• Tropical cascade darner, Spinaeschna watsoni

genus: Telephlebia

• Southern evening darner, Telephlebia brevicauda
• Northern evening darner, Telephlebia cyclops
• Eastern evening darner, Telephlebia godeffroyi
• Tropical evening darner, Telephlebia tillyardi
• Coastal evening darner, Telephlebia tryoni
• Carnarvon evening darner, Telephlebia undia

Austropetaliidae

genus: Archipetalia

• Tasmanian redspot, Archipetalia auriculata (Tasmania)

genus: Austropetalia

• Northern redspot, Austropetalia annaliese (Eastern New South Wales)
• Waterfall redspot, Austropetalia patricia (Coastal New South Wales)
• Alpine redspot, Austropetalia tonyana (Southern New South Wales and Victoria between 600–1800 metres elevation)

Corduliidae

genus: Hemicordulia

• Australian emerald, Hemicordulia australiae (Widespread in eastern and south-western Australia)
• Fat-bellied emerald, Hemicordulia continentalis (Coastal Queensland)
• Desert emerald, Hemicordulia flava (Central Australia)
• Yellow-spotted emerald, Hemicordulia intermedia (Northern Australia)
• Slender emerald, Hemicordulia kalliste (Northern Australia)
• Pilbara emerald, Hemicordulia koomina (Pilbara, Western Australia)
• Superb emerald, Hemicordulia superba (Northern New South Wales and south-eastern Queensland)
• Tau emerald, Hemicordulia tau (Widespread across Australia)

genus: Pentathemis

• Metallic tigerhawk, Pentathemis membranulata (Northern Australia)

genus: Procordulia

• Western swamp emerald, Procordulia affinis (South-western Western Australia)
• Eastern swamp emerald, Procordulia jacksoniensis (Eastern New South Wales, Victoria, south-eastern South Australia, Tasmania)

Gomphidae

genus: Antipodogomphus

• Southern dragon, Antipodogomphus acolythus Queensland except for Cape York and south-west, New South Wales except south-east and Victoria
• Top End dragon, Antipodogomphus dentosus Top End of Northern Territory
• Cape York dragon, Antipodogomphus edentulus Cape York, Queensland
• Pilbara dragon, Antipodogomphus hodgkini Pilbara region of Western Australia
• Northern dragon, Antipodogomphus neophytus Northern Australia, from Western Australia to Queensland
• Spinehead dragon, Antipodogomphus proselythus Coastal Queensland

genus: Armagomphus

• Armourtail, Armagomphus armiger South-western Western Australia.

Austrogomphus guerini

genus: Austroepigomphus

subgenus Austroepigomphus

• Twinspot hunter, Austroepigomphus praeruptus (South-eastern South Australia, Victoria)

subgenus Xerogomphus

• Western red hunter, Austrogomphus gordoni (Western Australia except the Kimberleys and central Australia)
• Flame-tipped hunter, Austrogomphus turneri (Northern Australia, from Western Australia to Queensland)

genus: Austrogomphus

subgenus Austrogomphus

• Murray River hunter, Austrogomphus angelorum (South-eastern South Australia, Victoria, western New South Wales)
• Toothed hunter, Austrogomphus arbustorum (Northern Queensland)
• Inland hunter, Austrogomphus australis (Victoria, south-eastern South Australia, southern Queensland, north-eastern and western New South Wales)
• Western inland hunter, Austrogomphus collaris (South-western Western Australia)
• Unicorn hunter, Austrogomphus cornutus (Victoria, eastern New South Wales, mid- and southern Queensland)
• Northern river hunter, Austrogomphus doddi (North-eastern Queensland)
• Yellow-striped hunter, Austrogomphus guerini (South-eastern Queensland, eastern New South Wales, Victoria, south-eastern South Australia, Tasmania)
• Pimple-headed hunter, Austrogomphus mjobergi (Northern Northern Territory, north-western Western Australia, north-eastern Queensland)
• Kimberly hunter, Austrogomphus mouldsorum (North-western Western Australia)
• Jade hunter, Austrogomphus ochraceus (South-eastern Queensland, eastern New South Wales, southern Victoria)
• Tiny hunter, Austrogomphus pusillus (North-western Western Australia)

subgenus Pleiogomphus

• Pale hunter, Austrogomphus amphiclitus (Eastern New South Wales, eastern and south-western Queensland)
• Dark hunter, Austrogomphus bifurcatus (North-eastern Queensland)
• Fork hunter, Austrogomphus divaricatus (Eastern Queensland)
• Lemon-tipped hunter, Austrogomphus prasinus (North-eastern Queensland)

genus: Hemigomphus

• Black vicetail, Hemigomphus atratus Mid coastal Queensland
• Zebra vicetail, Hemigomphus comitatus Mid and north coastal Queensland
• Wallum vicetail, Hemigomphus cooloola Coastal New South Wales
• Southern vicetail, Hemigomphus gouldii Coastal South Australia, Victoria, New South Wales, southern Queensland
• Stout vicetail, Hemigomphus heteroclytus Coastal South Australia, Victoria, New South Wales, southern Queensland
• Kakadu vicetail, Hemigomphus magela Northern Territory
• Rainforest vicetail, Hemigomphus theischingeri Mid and northern coastal Queensland

genus: Ictinogomphus

• Australian tiger, Ictinogomphus australis
• Pilbara tiger, Ictinogomphus dobsoni
• Cape York tiger, Ictinogomphus paulini

genus: Odontogomphus

• Pinchtail, Odontogomphus donnellyi (Northern coastal Queensland)

genus: Zephyrogomphus

• Lilac hunter, Zephyrogomphus lateralis (South-western Western Australia)
• Rainforest hunter, Zephyrogomphus longipositor (North-eastern Queensland)

Libellulidae

genus: Aethriamanta

• Square-spot basker, Aethriamanta circumsignata (Northern Australia)
• L-spot basker, Aethriamanta nymphaeae (Northern Australia)

genus: Agrionoptera

• Red swampdragon, Agrionoptera insignis allogenes (Northern New South Wales, eastern Queensland, Northern Territory)
• Striped swampdragon, Agrionoptera longitudinalis biserialis (North-eastern Queensland)

genus: Austrothemis

• Swamp flat-tail, Austrothemis nigrescens (Southern Australia)

genus: Brachydiplax

• Palemouth, Brachydiplax denticauda
• Darkmouth, Brachydiplax duivenbodei

genus: Camacinia

• Black knight, Camacinia othello

genus: Crocothemis

• Black-headed skimmer, Crocothemis nigrifrons

genus: Diplacodes

• Wandering percher, Diplacodes bipunctata (Widespread across Australia)
• Scarlet percher, Diplacodes haematodes (Widespread across Australia)
• Black-faced percher, Diplacodes melanopsis (Coastal southern Queensland, coastal New South Wales, Victoria)
• Charcoal-winged percher, Diplacodes nebulosa (Northern Australia)
• Chalky percher, Diplacodes trivialis (Northern Australia)

genus: Huonia

• Forestwatcher, Huonia melvillensis

genus: Hydrobasileus

• Water prince, Hydrobasileus brevistylus

genus: Lathrecista

• Australasian slimwing, Lathrecista asiatica festa

genus: Macrodiplax

• Wandering pennant, Macrodiplax cora

genus: Nannodiplax

• Pygmy percher, Nannodiplax rubra

genus: Nannophlebia

• Elusive archtail, Nannophlebia eludens
• Pilbara archtail, Nannophlebia injibandi
• Top End archtail, Nannophlebia mudginberri
• Common archtail, Nannophlebia risi

genus: Nannophya

• Australian pygmyfly, Nannophya australis
• Eastern pygmyfly, Nannophya dalei
• Artesian pygmyfly, Nannophya fenshami
• Western pygmyfly, Nannophya occidentalis
• Scarlet pygmyfly, Nannophya paulsoni

genus: Neurothemis

• Spotted grasshawk, Neurothemis oligoneura
• Painted grasshawk, Neurothemis stigmatizans

genus: Notolibellula

• Bicoloured skimmer, Notolibellula bicolor

genus: Orthetrum

• Speckled skimmer, Orthetrum balteatum
• Brownwater skimmer, Orthetrum boumiera
• Blue skimmer, Orthetrum caledonicum
• Rosy skimmer, Orthetrum migratum
• Slender skimmer, Orthetrum sabina
• Green skimmer, Orthetrum serapia
• Fiery skimmer, Orthetrum villosovittatum

genus: Pantala

• Wandering glider, Pantala flavescens

genus: Potamarcha

• Swampwatcher, Potamarcha congener

genus: Raphismia

• Spiny-chested percher, Raphismia bispina

genus: Rhodothemis

• Red arrow, Rhodothemis lieftincki

genus: Rhyothemis

• Iridescent flutterer, Rhyothemis braganza
• Graphic flutterer, Rhyothemis graphiptera
• Yellow-striped flutterer, Rhyothemis phyllis
• Sapphire flutterer, Rhyothemis princeps
• Jewel flutterer, Rhyothemis resplendens

genus: Tetrathemis

• Rainforest elf, Tetrathemis irregularis

genus: Tholymis

• Twister, Tholymis tillarga

genus: Tramea

Male common glider - Darwin

• Dune glider, Tramea eurybia
• Common glider, Tramea loewii
• Northern glider, Tramea transmarina
• Narrow-lobed glider, Tramea stenoloba

genus: Urothemis

• Red baron, Urothemis aliena

genus: Zyxomma

• Short-tailed duskdarter, Zyxomma elgneri
• Large duskdarter, Zyxomma multinervorum
• Long-tailed duskdarter, Zyxomma petiolatum

Macromiidae

genus: Macromia

• Australian cruiser, Macromia tillyardi
• Rainforest cruiser, Macromia viridescens

Petaluridae

genus: Petalura

• South-eastern petaltail, Petalura gigantea
• Western petaltail, Petalura hesperia
• Giant petaltail, Petalura ingentissima
• Coastal petaltail, Petalura litorea
• Beautiful petaltail, Petalura pulcherrima

Synthemistidae

genus: Archaeosynthemis

• Twinspot tigertail, Archaeosynthemis leachii
• Western brown tigertail, Archaeosynthemis occidentalis
• Eastern brown tigertail, Archaeosynthemis orientalis
• Spiny tigertail, Archaeosynthemis spiniger

genus: Austrosynthemis

• Turquoise tigertail, Austrosynthemis cyanitincta

genus: Choristhemis

• Yellow-tipped tigertail, Choristhemis flavoterminata
• Delicate tigertail, Choristhemis olivei

genus: Eusynthemis

• Variable tigertail, Eusynthemis aurolineata
• Mount Lewis tigertail, Eusynthemis barbarae
• Small tigertail, Eusynthemis brevistyla
• Cooloola tigertail, Eusynthemis cooloola
• Carnarvon tigertail, Eusynthemis deniseae
• Southern tigertail, Eusynthemis guttata
• Pretty tigertail, Eusynthemis netta
• Black tigertail, Eusynthemis nigra
• Swift tigertail, Eusynthemis rentziana
• Rainforest tigertail, Eusynthemis tenera
• Mountain tigertail, Eusynthemis tillyardi
• Barrington tigertail, Eusynthemis ursa
• Beech tigertail, Eusynthemis ursula
• Golden tigertail, Eusynthemis virgula

genus: Parasynthemis

• Royal tigertail, Parasynthemis regina

genus: Synthemiopsis

• Tasmanian spotwing, Synthemiopsis gomphomacromioides

genus: Synthemis

• Swamp tigertail, Synthemis eustalacta
• Tasmanian swamp tigertail, Synthemis tasmanica

genus: Tonyosynthemis

• Clavicle tigertail, Tonyosynthemis claviculata
• Slender tigertail, Tonyosynthemis ofarrelli

Superfamily Libelluloidea

genus: Apocordulia

• Nighthawk, Apocordulia macrops

genus: Archaeophya

• Horned urfly, Archaeophya adamsi
• Magnificent urfly, Archaeophya magnifica

genus: Austrocordulia

• Sydney hawk, Austrocordulia leonardi
• Eastern hawk, Austrocordulia refracta
• Top End hawk, Austrocordulia territoria

genus: Austrophya

• Summit mystic, Austrophya monteithorum
• Rainforest mystic, Austrophya mystica

genus: Cordulephya

• Common shutwing, Cordulephya pygmaea
• Tropical shutwing, Cordulephya bidens
• Clubbed shutwing, Cordulephya divergens
• Mountain shutwing, Cordulephya montana

genus: Hesperocordulia

• Orange streamcruiser, Hesperocordulia berthoudi

genus: Lathrocordulia

• Queensland swiftwing, Lathrocordulia garrisoni
• Western swiftwing, Lathrocordulia metallica

genus: Micromidia

• Forest mosquitohawk, Micromidia atrifrons
• Early mosquitohawk, Micromidia convergens
• Thursday Island mosquitohawk, Micromidia rodericki

genus: Pseudocordulia

• Circle-tipped mistfly, Pseudocordulia circularis
• Ellipse-tipped mistfly, Pseudocordulia elliptica

References

• "Order ODONATA". Australian Faunal Directory. Australian Biological Resources Study. 2022. Retrieved 1 September 2024.
• "Theischinger, Günther; Hawking, John (2021). The Complete Field Guide to Dragonflies of Australia (2nd ed.). Melbourne, Australia: CSIRO Publishing. p. 406. ISBN 9781486313747.
• Watson, J.A.L.; Theischinger, G.; Abbey, H.M. (1991). The Australian Dragonflies: A Guide to the Identification, Distributions and Habitats of Australian Odonata. Melbourne: CSIRO. ISBN 0643051368.
• Theischinger, Gunther; Endersby, Ian (2009). Identification Guide to the Australian Odonata (PDF). Department of Environment, Climate Change and Water NSW. p. 241. ISBN 978 1 74232 475 3.
• Paulson, D.; Schorr, M.; Abbott, J.; Bota-Sierra, C.; Deliry, C.; Dijkstra, K.-D.; Lozano, F. (2024). "World Odonata List". OdonataCentral, University of Alabama.