Ecosemiotics is a branch of semiotics in its intersection with human ecology, ecological anthropology and ecocriticism. It studies sign processes in culture, which relate to other living beings, communities, and landscapes. Ecosemiotics also deals with sign-mediated aspects of ecosystems. 
For ecosemiotics, environment has semiotic quality in different ways and levels. Material environment has affordances and potentials to participate in sign relations. Animal species attribute meanings to the environment based on their needs and umwelts. In human culture, environment can become meaningful in literary and artistic representations or through symbolization of animals or landscapes. Cultural representations of the environment in turn influence the natural environment through human actions.
Ecosemiotics analyzes processes, transmissions and problems that occur in and between the different semiotic layers of the environment. The central focus of ecosemiotics concerns the role of concepts (sign-based models people have) in designing and changing the environment. Concepts of ecosemiotic analysis are, for instance, semiocide, affordance, ecofield, consortium, dissent.
- Nöth, Winfried 1998. Ecosemiotics. Sign Systems Studies 26: 332–343.
- Kull, Kalevi 1998. Semiotic ecology: Different natures in the semiosphere. Sign Systems Studies 26: 344–371.
- Maran, Timo; Kull, Kalevi 2014. Ecosemiotics: main principles and current developments. Geografiska Annaler: Series B, Human Geography 96(1): 41–50.
- Siewers, Alfred Kentigern 2013. Re-Imagining Nature: Environmental Humanities and Ecosemiotics. Bucknell University Press.
- Farina, Almo; Santolini, Riccardo; Pagliaro, Giacomo; Scozzafava, Silvia; Schipani, Ileana 2005. Ecosemiotics: A new field of competence for ecology to overcome the frontier between environmental complexity and human culture in the Mediterranean. Israel Journal of Plant Sciences 53(3/4): 167–175.
- Nielsen, Søren Nors 2007. Towards an ecosystem semiotics: Some basic aspects for a new research programme. Ecological Complexity 4(3): 93–101.
- *Maran, Timo 2007. Towards an integrated methodology of ecosemiotics: The concept of nature-text. Sign Systems Studies 35(1/2): 269–294.
- Lindström, Kati; Kull, Kalevi; Palang, Hannes 2014. Landscape semiotics: Contribution to culture theory. In: Lang, Valter; Kull, Kalevi (eds.), Estonian Approaches to Culture Theory. Tartu: University of Tartu Press, 110–132.