Die Gattung Nepenthes

      The first pages of each of the four parts of "Die Gattung Nepenthes" from the March, April, May and June issues of Wiener Illustrirte Garten-Zeitung
      The three illustrations published in Beck's monograph, showing various pitchers drawn from cultivated plants.
      Left: N. curtisii var. superba (N. maxima) and N. × mixta
      Centre: N. curtisii (N. maxima), N. northiana, N. rafflesiana, N. rajah, N. veitchii, N. × dominii, N. × hookeriana, and N. × intermedia
      Right: N. albomarginata, N. ampullaria, N. bicalcarata, N. distillatoria, N. gracilis, N. × mastersiana, and N. × williamsii

      "Die Gattung Nepenthes: Eine monographische Skizze" (lit. "The genus Nepenthes: A monographic sketch") is a German-language monograph by Günther Beck von Mannagetta und Lerchenau on the tropical pitcher plants of the genus Nepenthes.[1] It was published in 1895 in four parts, spread over the March, April, May and June issues of Wiener Illustrirte Garten-Zeitung.[1]

      Content

      Beck recognised and enumerated 67 major taxa, almost half of which were hybrids. Additionally, seven taxa and unnamed horticultural hybrids are listed at the end of the work under "Ungenügend bekannte Arten, Mischarten" (insufficiently known species, mixed species).[1]

      Beck described four new species: N. fallax (later synonymised with N. stenophylla, although this is disputed),[2]N. hispida, N. smithii (later synonymised with N. distillatoria),[2] and N. spuria (later synonymised with N. northiana).[2] Beck was also the first to publish N. sumatrana under its present binomial combination,[3][4] although he introduced it under the entry for N. maxima with the words "Hierzu gehört als Varietät: N. sumatrana" (this includes a variety: N. sumatrana).[1]

      Beck also introduced a number of new varieties. These included the species varieties N. albomarginata var. tomentella, N. albomarginata var. typica, N. gracilis var. longinodis, N. gracilis var. teysmanniana, N. rafflesiana var. ambigua, N. rafflesiana var. excelsior (later synonym of N. × hookeriana), and N. rafflesiana var. typica, as well as the hybrid variety N. hybrida var. typica (N. gracilis × N. khasiana).[4]

      A number of taxon names that Beck did not recognise as valid nonetheless appeared in print for the first time in his monograph. These included the species synonyms N. edgeworthii (synonym of N. edwardsiana) and N. speciosa (synonym of N. distillatoria), as well as the hybrid synonyms N. anerleyense and N. arnoldiense (both synonyms of N. × stewartii).[4] While the name N. hookerae had first appeared in the literature four years prior to Beck's work, his concept of this hybrid involved a more complex cross (N. mirabilis × (N. rafflesiana × N. ampullaria)); it is considered a later synonym of N. × stewartii.[4]

      Beck was the first to unite N. edwardsiana with N. villosa,[5] considering the former a variety or form of the latter.[1] This synonymy stood until John Muirhead Macfarlane's 1908 monograph, "Nepenthaceae", and the species are considered distinct today.[2] Beck thought that N. burkei likely represented a form of N. boschiana ("Hierzu gehört offenbar auch als Form: Nepenthes Burkeii").[1] He likewise considered N. singalana to be a form of N. sanguinea ("Kaum als Form von N. sanguinea abzutrennen ist: Nepenthes singalana").[1]

      Species

      The following 67 taxa are enumerated and detailed in "Die Gattung Nepenthes". Only taxa considered hybrids by Beck are indicated here as such (therefore the natural hybrid N. × trichocarpa is shown as a species). Taxon names are listed as they appear in Beck's monograph, including orthographic variants, though specific epithets derived from proper nouns have been decapitalised.

      1. N. alata
      2. N. albomarginata
      3. N. × amesiana
      4. N. ampullaria
      5. N. × atrosanguinea
      6. N. bicalcarata
      7. N. bongso
      8. N. boschiana
      9. N. celebica
      10. N. × chelsoni
      11. N. × cincta
      12. N. × coccinea
      13. N. × compacta
      14. N. × courtii
      15. N. × cylindrica
      16. N. destillatoria [sic]
      17. N. × dicksoniana
      18. N. × dominii
      19. N. × dormanniana
      20. N. echinostoma
      21. N. × edinensis
      22. N. eustachya
      23. N. fallax
      24. N. gracilis
      25. N. × henryana
      26. N. × hibberdii
      27. N. hirsuta
      28. N. hispida
      29. N. × hookerae
      30. N. × hybrida
      31. N. × intermedia
      32. N. khasiana
      33. N. × lawrenciana
      34. N. lowii
      35. N. macrostachya
      36. N. madagascariensis
      37. N. × mastersiana
      38. N. maxima
      39. N. melamphora
      40. N. × mixta
      41. N. × morganiae
      42. N. northiana
      43. N. × outramiana
      44. N. × paradisae
      45. N. pervillei
      46. N. rafflesiana
      47. N. rajah
      48. N. × ratcliffiana
      49. N. reinwardtiana
      50. N. × robusta
      51. N. × rubro maculata
      52. N. × rufescens
      53. N. sanguinea
      54. N. × sedeni
      55. N. smithii
      56.  ? N. spuria
      57. N. stenophylla
      58. N. × stewartii
      59. N. × superba
      60. N. tentaculata
      61. N. trichocarpa
      62. N. veitchii
      63. N. ventricosa
      64. N. vieillardii
      65. N. villosa
      66. N. × williamsii
      67. N. × wrigleyana
      Ungenügend bekannte Arten, Mischarten
      1. N. bernaysii
      2. N. burbidgeae
      3. N. cristata
      4. N. × lyrata
      5. N. loddigesi
      6. N. veitchii × N. rafflesiana
      7. N. sanguinea × N. destillatoria

      Infrageneric classification

      The infrageneric classification in "Die Gattung Nepenthes" follows that introduced by Joseph Dalton Hooker in his 1873 monograph, "Nepenthaceae". Hooker placed N. pervillei in the monotypic Anourosperma, distinguishing it on the basis of its round seeds, and subsumed all remaining species in the second section, Eunepenthes.[6] Beck kept the two sections created by Hooker (spelling Anourosperma as Anowrosperma), but divided Eunepenthes into three subgroups: Apruinosae, Pruinosae, and Retiferae, the last containing only N. lowii.[1]

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      References

      1. ^ a b c d e f g h (German) Beck, G. 1895. Die Gattung Nepenthes. Wiener Illustrirte Garten-Zeitung 20(3–6): 96–107, 141–150, 182–192, 217–229.
      2. ^ a b c d McPherson, S.R. 2009. Pitcher Plants of the Old World. 2 volumes. Redfern Natural History Productions, Poole.
      3. ^ Clarke, C.M. 2001. Nepenthes of Sumatra and Peninsular Malaysia. Natural History Publications (Borneo), Kota Kinabalu.
      4. ^ a b c d Schlauer, J. N.d. Query results: Wien.Ill.Gart.Z.. Carnivorous Plant Database.
      5. ^ Danser, B.H. 1928. The Nepenthaceae of the Netherlands Indies. Bulletin du Jardin Botanique de Buitenzorg, Série III, 9(3–4): 249–438.
      6. ^ Hooker, J.D. 1873. Nepenthaceae. In: A. de Candolle Prodromus Systematis Naturalis Regni Vegetabilis 17: 90–105.
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      Last modified on 21 March 2013, at 21:46