Cystodium sorbifolium

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Cystodium is a fern in its own family, Cystodiaceae.[2] It contains a single species: Cystodium sorbifolium (Sm.) J.Sm. 1841.[3] Because it looks like a small tree fern, it had previously been placed in the tree fern family Dicksoniaceae. Subsequent analysis had moved it to the Lindsaeaceae,[3] but the most recent phylogenetic studies have placed it in its own separate family, Cystodiaceae, with a sister relationship to the current Lindsaeaceae.[4] A fossil species of the genus Cystodium sorbifolioides is known from the Cenomanian aged Burmese amber in Myanmar.[5]

Cystodium sorbifolium
Scientific classification edit
Kingdom: Plantae
Clade: Tracheophytes
Division: Polypodiophyta
Class: Polypodiopsida
Order: Polypodiales
Suborder: Lindsaeineae
Family: Cystodiaceae
J.R.Croft 1986
Genus: Cystodium
J.Sm. 1841
C. sorbifolium
Binomial name
Cystodium sorbifolium
(Sm.) J.Sm.

Cystodiopteris Rauschert 1982[1]


Cystodium is distributed through lowland rainforests from Borneo to New Guinea and nearby islands, as well as the Solomon Islands.[3]


  1. ^ Cystodiopteris USDA-ARS Germplasm Resources Information Network (GRIN) 28 January 2010
  2. ^ Christenhusz, Maarten; Xian-Chun Zhang & Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns" (PDF). Phytotaxa. 19 (18 Feb. 2011): 7–54. doi:10.11646/phytotaxa.19.1.2.
  3. ^ a b c Korall, Petra; Conant, David S.; Schneider, Harald; Ueda, Kunihiko; Nishida, Harufumi; Pryer, Kathleen M. (2006). "On the Phylogenetic Position of Cystodium: It's Not a Tree Fern – It's a Polypod!" (PDF). American Fern Journal. 96 (2): 45–53. doi:10.1640/0002-8444(2006)96[45:OTPPOC]2.0.CO;2. Archived from the original (PDF) on 2008-08-20. Retrieved 2012-02-02.
  4. ^ Lehtonen, Samuli; Tuomisto, Hanna; Rouhan, Germinal; Christenhusz, Maarten J. M. (2010). "Phylogenetics and classification of the pantropical fern family Lindsaeaceae". Botanical Journal of the Linnean Society. 163 (3): 305–359. doi:10.1111/j.1095-8339.2010.01063.x.
  5. ^ Regalado, Ledis; Schmidt, Alexander R.; Appelhans, Marc S.; Ilsemann, Bork; Schneider, Harald; Krings, Michael; Heinrichs, Jochen (December 2017). "A fossil species of the enigmatic early polypod fern genus Cystodium (Cystodiaceae) in Cretaceous amber from Myanmar". Scientific Reports. 7 (1): 14615. doi:10.1038/s41598-017-14985-7. ISSN 2045-2322. PMC 5668433. PMID 29097728.