Pterostichinae

Pterostichinae is a subfamily of ground beetles (family Carabidae). It belongs to the advanced harpaline assemblage, and if these are circumscribed sensu lato as a single subfamily, Pterostichinae are downranked to a tribe Pterostichini. However, as the former Pterostichitae supertribe of the Harpalinae as loosely circumscribed does seem to constitute a lineage rather distinct from Harpalus, its core group is here considered to be the present subfamily and the Harpalinae are defined more narrowly.

Pterostichinae
Image of the common Eurasian species Poecilus cupreus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Family:	Carabidae
Subfamily:	Pterostichinae Bonelli, 1810

They are usually mid-sized and rather stout ground beetles. Coloration is typically dark and without conspicuous patterns, but often with a strong sheen like polished metal. They are widely distributed and inhabit a wide range of terrestrial habitats. Unlike the more basal ground beetles which only eat small animals, the Pterostichinae include a large proportion of omnivorous or even herbivorous taxa.

Systematics

This group includes the following tribes and genera:

Tribe Chaetodactylini Tschitscherine, 1903

• Chaetodactyla Tschitscherine, 1897

Tribe Cnemalobini Germain, 1911

• Cnemalobus Guerin-Meneville, 1838

Tribe Cratocerini Lacordaire, 1854

• Abacaelostus Straneo, 1952
• Andrewesinulus Straneo, 1938
• Apsidocnemus Alluaud, 1936
• Barylaus Liebherr, 1985
• Brachidius Chaudoir, 1852
• Caecocaelus Straneo, 1949
• Caelostomus W.S.MacLeay, 1825
• Camptogenys Tschitscherine, 1899
• Capabatus Csiki, 1930
• Catapiesis Solier, 1835
• Cratocerus Dejean, 1829
• Crenulostrigus Straneo, 1942
• Cyrtolaus Bates, 1882
• Dactyleurys Tschitscherine, 1899
• Dactylinius Straneo, 1941
• Diachipteryx Alluaud, 1925
• Diceromerus Chaudoir, 1873
• Dromistomus Jeannel, 1948
• Drymonaxus Straneo, 1941
• Feostoma Straneo, 1941
• Hemitelestus Alluaud, 1895
• Homalomorpha Brullé, 1837
• Hoplizomenus Chaudoir, 1873
• Leleuporites Straneo, 1960
• Madapelmus Dajoz, 1985
• Monodryxus Straneo, 1942
• Oxyglychus Straneo, 1938
• Pachycaecus Straneo, 1971
• Pachyroxochus Straneo, 1942
• Platyxythrius Lorenz, 1998
• Stegazopteryx Will, 2004
• Stomonaxellus Tschitscherine, 1901
• Strigomerodes Straneo, 1939
• Strigomerus Chaudoir, 1873
• Trichillinus Straneo, 1938

Tribe Microcheilini Jeannel, 1948

• Microcheila Brulle, 1834

Tribe Morionini Brulle, 1835

• Buderes Murray, 1857
• Hyperectenus Alluaud, 1935
• Hyperion Castelnau, 1834
• Megamorio Chaudoir, 1880
• Morion Latreille, 1810
• Morionidius Chaudoir, 1880
• Moriosomus Motschulsky, 1864
• Platynodes Westwood, 1846
• Stereostoma Murray, 1857

Tribe Pterostichini Bonelli, 1810

• (See Pterostichini for ~180 genera)

Tribe Zabrini Bonelli, 1810

• (See Zabrini for 3 genera)