Root hair

Root hairs, or absorbent hairs, are tubular outgrowths of a trichoblast, a hair-forming cell on the epidermis of a plant root. These structures are lateral extensions of a single cell and are only rarely branched. They are found in the region of maturation, also called the zone of differentiation[1] of the root. Just prior to and during root hair cell development, there is elevated phosphorylase activity.[2] Plants absorb water through the roots from the soil by bulk flow[citation needed]. Root hair cells are adapted to this process by increasing root surface area for the purpose of taking in more water. The large vacuole inside root hair cells makes this intake much more efficient[citation needed].

Drawing of root tip, showing young root hairs

FunctionEdit

Most water absorption happens in the root hairs. The root hairs are long which allows them to penetrate between soil particles, and prevent harmful bacterial organisms from entering the plant through the xylem vessels[citation needed]. Increasing the surface area of these hairs allows plants to be more efficient in absorbing nutrients and establish relationships with microbes.[3] Cross-section of root hair cell: a roughly rectangular shape with a long, thin tail extending to the right and a nucleus at the top left. This happens because the water in the soil has a higher water potential than the cytoplasm of the root hairs. The function of root hairs is to collect water and mineral nutrients that are present in the soil and take this solution up through the roots to the rest of the plant. As root hair cells do not carry out photosynthesis, they do not contain chloroplasts.

FormationEdit

Root hair cells are outgrowths at a tip of the plant's roots. Root hair cells vary between 15 and 17 micrometres in diameter, and 80 to 1,500 micrometres in length.[4] They are found only in the zone of maturation, and not the zone of elongation, possibly because any root hairs that arise are sheared off as the root elongates and moves through the soil.[citation needed] Root hairs grow quickly, at least 1μm/min, making them particularly useful for research on cell expansion.[5]

ImportanceEdit

Root hairs form an important surface as they are needed to absorb most of the water and nutrients needed for the plant. They are also directly involved in the formation of root nodules in legume plants. The root hairs curl around the bacteria which allows for the formation of an infection thread through into the dividing cortical cells to form the nodule.[6]

Having a large surface area, the active uptake of water and minerals through root hairs is highly efficient. Root hair cells also secrete acid (H+ from malic acid) which exchanges and helps solubilize the minerals into ionic form, making the ions easier to absorb.[7]

SurvivalEdit

This way, the root hair coverage stays the same. When a new root hair cell grows, it excretes a hormone so that the other cells in close proximity to it are unable to grow one of these hairs. This ensures equal and efficient distribution of the actual hairs on these cells.[citation needed]

The act of re-potting or transplanting a plant can result in root hair cells being pulled off, perhaps to a significant extent, and such plants may therefore wilt for some time as a result.

See alsoEdit

ReferencesEdit

  1. ^ https://www.colby.edu/biology/BI237/roots.pdf
  2. ^ Dosier, Larry W.; Riopel, J. L. (1977). "Differential Enzyme Activity During Trichoblast Differentiation in Elodea Canadensis". American Journal of Botany. 64 (9): 1049–1056. doi:10.1002/j.1537-2197.1977.tb10794.x.
  3. ^ Grierson, C.; Schiefelbein, J. (2002). "Root Hairs". The Arabidopsis Book. 1: e0060. doi:10.1199/tab.0060. PMC 3243358. PMID 22303213.
  4. ^ Dittmar, cited in Esau, 1965
  5. ^ Grierson, Claire; Schiefelbein, John (1 January 2002). "Root Hairs". The Arabidopsis Book. 1: e0060. doi:10.1199/tab.0060. PMC 3243358. PMID 22303213.
  6. ^ Mergaert, Peter; Uchiumi, Toshiki; Alunni, Benoît; Evanno, Gwénaëlle; Cheron, Angélique; Catrice, Olivier; Mausset, Anne-Elisabeth; Barloy-Hubler, Frédérique; Galibert, Francis; Kondorosi, Adam; Kondorosi, Eva (28 March 2006). "Eukaryotic control on bacterial cell cycle and differentiation in the Rhizobium–legume symbiosis". Proceedings of the National Academy of Sciences. 103 (13): 5230–5235. Bibcode:2006PNAS..103.5230M. doi:10.1073/pnas.0600912103. PMC 1458823. PMID 16547129.
  7. ^ Gerke, Jörg; Römer, Wilhelm; Jungk, Albrecht (1994). "The excretion of citric and malic acid by proteoid roots ofLupinus albus L.; effects on soil solution concentrations of phosphate, iron, and aluminum in the proteoid rhizosphere in samples of an oxisol and a luvisol". Zeitschrift für Pflanzenernährung und Bodenkunde. 157 (4): 289–294. doi:10.1002/jpln.19941570408.

Further readingEdit