Protungulatum donnae

Protungulatum donnae is the type species for the genus Protungulatum, an extinct early form of eutherian mammal. Though it is by no means the earliest mammal in the fossil record, a 2013 study considers P. donnae to be the oldest undisputed placental mammal fossil,[1][2] though more recent examinations conclude that it was a more basal eutherian and that no placental predates the Paleocene.[3]

Temporal range: Late Cretaceous - Paleocene, Maastrichtian–Danian Possible Campanian record
Protungulatum donnae.JPG
Scientific classification

Sloan and Van Valen, 1965
P. donnae


  1. ^ O'Leary, Maureen A.; Bloch, Jonathan I.; Flynn, John J.; Gaudin, Timothy J.; Giallombardo, Andres; Giannini, Norberto P.; Goldberg, Suzann L.; Kraatz, Brian P.; Luo, Zhe-Xi; Meng, Jin; Novacek, Michael J.; Perini, Fernando A.; Randall, Zachary S.; Rougier, Guillermo; Sargis, Eric J.; Silcox, Mary T.; Simmons, Nancy b.; Spaulding, Micelle; Velazco, Paul M.; Weksler, Marcelo; Wible, John r.; Cirranello, Andrea L. (8 February 2013). "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667. Bibcode:2013Sci...339..662O. doi:10.1126/science.1229237. PMID 23393258. S2CID 206544776.
  2. ^ Wilford, John Noble (7 February 2013). "Rat-Size Ancestor Said to Link Man and Beast". New York Times. Retrieved 9 February 2013.
  3. ^ Halliday, Thomas J. D. (2015). "Resolving the relationships of Paleocene placental mammals" (PDF). Biological Reviews. 92 (1): 521–550. doi:10.1111/brv.12242. PMC 6849585. PMID 28075073.