The Phaneropterinae, the sickle-bearing bush crickets or leaf katydids, are a subfamily of insects within the family Tettigoniidae.[2][3] Nearly 2,060 species in 85 genera throughout the world are known.[citation needed] They are also known as false katydids[4] or round-headed katydids.[2]

Phaneroptera nana female (7973618058).jpg
Female Phaneroptera nana
Isophya pyrenaea male (15031536651).jpg
Male Isophya pyrenaea
Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Orthoptera
Suborder: Ensifera
Family: Tettigoniidae
Subfamily: Phaneropterinae
Burmeister, 1838[1]
  • Amblycoryphae Scudder, 1897
  • Phaneropteridae Burmeister, 1838

The name Phaneropterinae is based upon the Old World genus Phaneroptera (type species P. falcata), meaning "visible wing";[5] this refers to the exposed tips of the inner wings seen in many species, although some genera, notably in the tribes Barbitistini and Odonturini have become brachypterous.

Adult Phaneropterinae species with eggs


The legs of individuals in this subfamily vary from genus to genus, but, as in nearly all Orthoptera, the posterior (rear) legs are adapted to leaping, and as such are always much longer than other legs.

The Phaneropterinae are largely arboreal in habitat.[2] The vast majority of species live in shrubs and trees, feeding on leaves and twigs.[2] Some species might potentially cause significant damage, though usually superficial, when present in large numbers, but this is rare; they usually are solitary, unlike meadow grasshoppers, so much so that they seldom come to human notice.[2]

The ovipositor and male genitalia vary according to the genus. The Phaneropterinae differ from other subfamilies of Tettigoniidae (and other Orthoptera) in their oviposition; their eggs are rarely deposited in the earth, but are either glued in double rows to twigs, or are inserted in the edges of leaves.[2]


The subfamily Phaneropterinae was first erected in 1838 by German zoologist Hermann Burmeister.[3] As of January 2019, Orthoptera Species File lists the following tribes, subtribes, genus groups, and genera.[3]


Auth.: Brunner von Wattenwyl, 1878 – Europe, the Middle East, Africa


Auth.: Brunner von Wattenwyl, 1878 – Americas, Africa


Auth.: Jacobson, 1905 – Europe to central Asia

Isophya species mating
  1. Ancistrura Uvarov, 1921 - monotypic Ancistrura nigrovittata (Brunner von Wattenwyl, 1878)
  2. Andreiniimon Capra, 1937 - monotypic Andreiniimon nuptialis (Karny, 1918)
  3. Barbitistes Charpentier, 1825
  4. Dasycercodes Bei-Bienko, 1951 - monotypic Dasycercodes iranicus Bey-Bienko, 1951
  5. Euconocercus Bei-Bienko, 1950
  6. Isoimon Bei-Bienko, 1954 - monotypic Isoimon riabovi (Uvarov, 1927)
  7. Isophya Brunner von Wattenwyl, 1878
  8. Kurdia Uvarov, 1916
  9. Leptophyes Fieber, 1853
  10. Metaplastes Ramme, 1939
  11. Orthocercodes Bei-Bienko, 1951 - monotypic Orthocercodes zarudnyi (Uvarov, 1930)
  12. Phonochorion Uvarov, 1916
  13. Poecilimon Fischer, 1853
  14. Polysarcus Fieber, 1853


Auth.: Massa, 2016 – Africa


Auth.: Brunner von Wattenwyl, 1878 – Africa, India, China, Indochina to Australia

Shirakisotima japonica


Dysonia sp.
Markia arizae

Auth.: Rehn, 1950 – tropical Americas


Auth.: Cadena-Castañeda, 2015 – tropical Americas


Auth.: Brunner von Wattenwyl, 1891 – Asia


Auth.: Brunner von Wattenwyl, 1878 – Africa, Asia-Pacific

Holochlora japonica female laying eggs


Auth.: Rehn, & Hebard, 1914 – tropical & subtropical Americas


Auth.: Massa, 2017 – eastern Africa


Auth.: Hebard, 1922 – Asia


Auth.: Brunner von Wattenwyl, 1878 – Americas

Eggs of Microcentrum.


Auth.: Brunner von Wattenwyl, 1878 – Asia


Auth.: Karsch, 1890 - central and western Africa


Auth.: Brunner von Wattenwyl, 1878 – Americas, Iberian peninsula, Africa, Pacific islands

Obolopteryx castanea


Auth.: Karsch, 1889 - Africa


Auth.: Brunner von Wattenwyl, 1878 - Africa


Auth.: Cadena-Castañeda, 2015 – South America


Auth.: Burmeister, 1838 – distributed worldwide

Phaneroptera nana nymph


Auth.: Karsch, 1889 – Africa


Auth.: Brunner von Wattenwyl, 1878


Auth.: Brunner von Wattenwyl, 1878 – South America


Auth.: Cadena-Castañeda, 2015 – Africa


Auth.: Brunner von Wattenwyl, 1878 – Africa


Auth.: Karsch, 1890 – Africa


Auth.: Cadena-Castañeda, 2014 – Central and South America


Auth.: Brunner von Wattenwyl, 1878 – Americas


Auth.: Brunner von Wattenwyl, 1878 – South America


Auth.: Brunner von Wattenwyl, 1878 – Africa, Sri Lanka


Auth.: Bei-Bienko, 1954 – Middle East, India, Malaysia


Auth.: Brunner von Wattenwyl, 1878 – Africa, Europe, Middle East, western Asia


Auth.: Cadena-Castañeda, 2015 – Africa, Asia

  • Vossiina Cadena-Castañeda, 2015 – Africa
  • Xantiina Cadena-Castañeda, 2015 – Asia
    • Paraxantia Liu & Kang, 2009 – China
    • Xantia Brunner von Wattenwyl, 1878 – Borneo


Zeuneria melanopeza

Auth.: Karsh, 1890 - tropical Africa

Genus groupsEdit

Subtribe Trachyzulphina Gorochov, 2014 – China, Indochina, Malesia

Genus groups include:

  • Acridopezae Brunner von Wattenwyl, 1878 – Australia
  • Aegimiae Brunner von Wattenwyl, 1878 – Central and South America
  • Aniarae Brunner von Wattenwyl, 1878 – South America, E. Africa
  • Centroferae Brunner von Wattenwyl, 1878 – South America
  • Cosmophylla Brunner von Wattenwyl, 1878 – South America, Madagascar, Australia
  • Dysmorphae Brunner von Wattenwyl, 1878 – Malesia
  • Ephippithytae Brunner von Wattenwyl, 1878 – Vietnam, Papua New Guinea, Australia
  • Eurypalpae Brunner von Wattenwyl, 1878 – Eastern China, Vietnam, Sumatra, Australia
  • Isopserae Brunner von Wattenwyl, 1878 – Eastern Africa, India, Indochina, Malesia
  • Karschiae Brunner von Wattenwyl, 1891
  • Leptoderae Brunner von Wattenwyl, 1878 – Sri Lanka, Malesia
  • Pseudophaneropterae Brunner von Wattenwyl, 1878 – Sri Lanka, Sumatra
  • ScambophyllaBrunner von Wattenwyl, 1878 – Indochina, Malesia
  • Scaphurae Westwood, 1838 – South America
  • Taeniomenae Brunner von Wattenwyl, 1878 – Australia
  • Turpiliae Brunner von Wattenwyl, 1878 – Central America, Caribbean

Incertae sedisEdit

These genera have not been placed in a tribe:


  1. ^ Burmeister H (1838) Handbuch der Entomologie 2 2(I-VIII):684
  2. ^ a b c d e f Blatchley WS. 1920. Orthoptera of northeastern America, Family VII. Tettigoniidae, Subfamily I, Phaneropterinae. Indianapolis, IN: Nature Publishing. 784 p. Phaneropterinae (pp. 459-494)
  3. ^ a b c Cigliano, M. M.; Braun, H.; Eades, D. C.; Otte, D. "subfamily Phaneropterinae Burmeister, 1838". Orthoptera Species File. Retrieved 4 January 2019.
  4. ^ Walker, Thomas J. "Singing Insects of North America (SINA), Phaneropterinae". Retrieved 2 January 2019.
  5. ^ Jaeger, Edmund Carroll (1959). A source-book of biological names and terms. Springfield, Ill: Thomas. ISBN 0-398-06179-3.

External linksEdit