Jaguar(Redirected from Panthera onca)
The jaguar (Panthera onca), is a wild cat species and the only extant member of the genus Panthera native to the Americas. The jaguar's present range extends from Southwestern United States and Mexico across much of Central America and south to Paraguay and northern Argentina. Though there are single cats now living within the western United States, the species has largely been extirpated from the United States since the early 20th century. It is listed as Near Threatened on the IUCN Red List; and its numbers are declining. Threats include loss and fragmentation of habitat.
Temporal range: 0.5–0 Ma Middle Pleistocene – Recent
|Male South American jaguar at Three Brothers River, São Paulo state, Brazil|
|Female at Piquiri River, Mato Grosso state, Brazil|
Red : Current range
Bright pink : Former range
The jaguar is the largest cat species in the Americas and the third-largest after the tiger and the lion. This spotted cat closely resembles the leopard, but is usually larger and sturdier. It ranges across a variety of forested and open terrains, but its preferred habitat is tropical and subtropical moist broadleaf forest, swamps and wooded regions. The jaguar enjoys swimming and is largely a solitary, opportunistic, stalk-and-ambush predator at the top of the food chain. As a keystone species it plays an important role in stabilizing ecosystems and regulating prey populations.
The jaguar has an exceptionally powerful bite, even relative to the other big cats. This allows it to pierce the shells of armored reptiles and to employ an unusual killing method: it bites directly through the skull of prey between the ears to deliver a fatal bite to the brain.
While international trade in jaguar or its body parts is prohibited, the cat is still frequently killed, particularly in conflicts with ranchers and farmers in South America. Although reduced, its range remains large. Given its historical distribution, the jaguar has featured prominently in the mythology of numerous indigenous American cultures, including those of the Maya and Aztec.
The word 'jaguar' is thought to derive from the Tupian word yaguara, meaning "beast of prey". The word entered English presumably via the Amazonian trade language Tupinambá, via Portuguese jaguar. The specific word for jaguar is yaguareté, with the suffix -eté meaning "real" or "true".
The word 'panther' derives from classical Latin panthēra, itself from the ancient Greek pánthēr (πάνθηρ). The Greek pan- (πάν), meaning "all", and thēr (θήρ), meaning "prey" bears the meaning of "predator of all animals". Use of the word for a beast originated in antiquity in the Orient, probably from India to Persia to Greece.
In Mexican Spanish, its nickname is el tigre: 16th century Spaniards had no native word in their language for the jaguar, which is smaller than a lion, but bigger than a leopard, nor had ever encountered it in the Old World, and so named it after the tiger, since its ferocity would have been known to them through Roman writings and popular literature during the Renaissance.
Onca is the Portuguese onça, with the cedilla dropped for typographical reasons, found in English as ounce for the snow leopard, Panthera uncia. It derives from the Latin lyncea lynx, with the letter L confused with the definite article (Italian lonza, Old French l'once).
The jaguar is the only extant New World member of the genus Panthera. Results of DNA analysis shows the lion, tiger, leopard, jaguar, snow leopard, and clouded leopard share a common ancestor, and that this group is between six and ten million years old; the fossil record points to the emergence of Panthera just two to 3.8 million years ago. The Panthera are thought to have evolved in Asia. The jaguar is thought to have diverged from a common ancestor of the Panthera species at least 1.5 million years ago and to have entered the American continent in the Early Pleistocene via Beringia, the land bridge that once spanned the Bering Strait. Results of jaguar mitochondrial DNA analysis indicates that the species' lineage evolved between 280,000 and 510,000 years ago.
Phylogenetic studies generally have shown the clouded leopard (Neofelis nebulosa) is basal to this group. The position of the remaining species varies between studies and is effectively unresolved.
Based on morphological evidence, British zoologist Reginald Innes Pocock concluded the jaguar is most closely related to the leopard. However, DNA evidence is inconclusive and the position of the jaguar relative to the other species varies between studies. Fossils of extinct Panthera species, such as the European jaguar (Panthera gombaszoegensis) and the American lion (Panthera atrox), show characteristics of both the lion and the jaguar.
In the 19th and 20th centuries, several jaguar type specimens formed the basis for descriptions of subspecies. In 1939, Reginald Innes Pocock recognized eight subspecies based on geographic origins and skull morphology of these specimens. Pocock did not have access to sufficient zoological specimens to critically evaluate their subspecific status, but expressed doubt about the status of several. Later consideration of his work suggested only three subspecies should be recognized. The description of P. o. palustris was based on a fossil skull. The author of Mammal Species of the World listed nine subspecies and both P. o. palustris or P. o. paraguensis separately.
Results of morphologic and genetic research indicate a clinal north–south variation between populations, but no evidence for subspecific differentiation. A subsequent, more detailed study confirmed the predicted population structure within jaguar populations in Colombia.
IUCN Red List assessors for the species and the Cat Classification Taskforce of the Cat Specialist Group do not recognize any jaguar subspecies as valid. The following table is based on the classification of the species provided in Mammal Species of the World.
|Formerly recognised subspecies||Region||Image|
|South America: Venezuela to the Amazon rainforest, coastal Peru, the Pantanal regions of Mato Grosso and Mato Grosso do Sul, Brazil, along the Paraguay River into Paraguay and northeastern Argentina|
|Central and North America: Colombia, Guatemala, Belize, El Salvador to Yucatán Peninsula, Mexico and southern Arizona, United States of America.|
The jaguar, a compact and well-muscled animal, is the largest cat in the New World and the largest carnivorous mammal in Central and South America. Size and weight vary considerably: weights are normally in the range of 56–96 kg (124–211 lb). Larger males have been recorded to weigh as much as 158 kg (348 lb) (roughly matching a tigress or lioness; however note this animal was weighed with a full stomach). The smallest females have low weights of 36 kg (79 lb). Females are typically 10–20 percent smaller than males. The length, from the nose to the base of the tail, of the cats varies from 1.12 to 1.85 m (3.7 to 6.1 ft). Their tails are the shortest of any big cat, at 45 to 75 cm (18 to 30 in) in length. Their legs are also short, considerably shorter when compared to a small tiger or lion in a similar weight range, but are thick and powerful. The jaguar stands 63 to 76 cm (25 to 30 in) tall at the shoulders. Compared to the similarly colored leopard, the jaguar is bigger, heavier and relatively stocky in build.
Further variations in size have been observed across regions and habitats, with size tending to increase from the north to south. A study of the jaguar in the Chamela-Cuixmala Biosphere Reserve on the Mexican Pacific coast, showed ranges of just about 50 kg (110 lb), about the size of a female cougar. Jaguars in Venezuela or Brazil are much larger with average weights of about 95 kg (220 lb) in males and of about 56 kilograms (123 lb) to 78 kilograms (172 lb) in females. In the Brazilian Pantanal, weights of 136 kilograms (300 lb) or more are not uncommon in old males, with the highest recorded weight, for a Jaguar weighed on an empty stomach being 148 kilograms (326 lb). Forest jaguars are frequently darker and considerably smaller than those found in open areas (the Pantanal is an open wetland basin), possibly due to the smaller numbers of large, herbivorous prey in forest areas.
A short and stocky limb structure makes the jaguar adept at climbing, crawling, and swimming. The head is robust and the jaw extremely powerful, it has the third highest bite force of all felids, after the tiger and lion. A 100 kg (220 lb) jaguar can bite with a force of 503.6 kgf (1110 lbf) at canine teeth and 705.8 kgf (1556 lbf) at carnassial notch. This strength adaptation allows the jaguar to pierce turtle shells. A comparative study of bite force adjusted for body size ranked it as the top felid, alongside the clouded leopard and ahead of the tiger and lion. It has been reported that "an individual jaguar can drag an 800 lb (360 kg) bull 25 ft (7.6 m) in its jaws and pulverize the heaviest bones". The jaguar hunts wild animals weighing up to 300 kg (660 lb) in dense jungle, and its short and sturdy physique is thus an adaptation to its prey and environment.
The base coat of the jaguar is generally a tawny yellow, but can range to reddish-brown, for most of the body. However, the ventral areas are white. The cat is covered in rosettes for camouflage in the dappled light of its forest habitat. The spots vary over individual coats and between individual jaguars: rosettes may include one or several dots, and the shapes of the dots vary. The spots on the head and neck are generally solid, as are those on the tail, where they may merge to form a band.
While the jaguar closely resembles the leopard, it is sturdier and heavier, and the two animals can be distinguished by their rosettes: the rosettes on a jaguar's coat are larger, fewer in number, usually darker, and have thicker lines and small spots in the middle that the leopard lacks. Jaguars also have rounder heads and shorter, stockier limbs compared to leopards.
The black morph is less common than the spotted form, but at about six percent of the population, it is several orders of magnitude above the mutation rate. Hence, it is being supported by selection. Some evidence indicates the melanism allele is dominant. The black form may be an example of heterozygote advantage; breeding in captivity is not yet conclusive on this. Melanistic jaguars (or “black” jaguars) occur primarily in parts of South America, and are virtually unknown in wild populations residing in the subtropical and temperate regions of North America; they have never been documented north of Mexico’s Isthmus of Tehuantepec.
Extremely rare albino individuals, sometimes called white panthers, also occur among jaguars, as with the other big cats. As usual with albinos in the wild, selection keeps the frequency close to the rate of mutation.
Distribution and habitat
At present, the jaguar's range extends from Mexico through Central America to South America, including much of Amazonian Brazil. The countries included in this range are Argentina, Belize, Bolivia, Colombia, Costa Rica (particularly on the Osa Peninsula), Ecuador, French Guiana, Guatemala, Guyana, Honduras, Nicaragua, Panama, Paraguay, Peru, Suriname, the United States and Venezuela. It is now locally extinct in El Salvador and Uruguay.
The jaguar has been an American cat since crossing the Bering Land Bridge during the Pleistocene epoch; the immediate ancestor of modern animals is Panthera onca augusta, which was larger than the contemporary cat. It occurs in the 400 km² Cockscomb Basin Wildlife Sanctuary in Belize, the 5,300 km² Sian Ka'an Biosphere Reserve in Mexico, the approximately 15,000 km2 Manú National Park in Peru, the approximately 26,000 km2 Xingu National Park in Brazil, and numerous other reserves throughout its range.
The inclusion of the United States in the list is based on occasional sightings in the southwest, particularly in Arizona, New Mexico and Texas. In the early 20th century, the jaguar's range extended as far north as the Grand Canyon, and as far west as Southern California. The jaguar is a protected species in the United States under the Endangered Species Act, which has stopped the shooting of the animal for its pelt. In 1996 and from 2004 on, hunting guides and wildlife officials in Arizona photographed and documented jaguars in the southern part of the state. Between 2004 and 2007, two or three jaguars have been reported by researchers around Buenos Aires National Wildlife Refuge in southern Arizona. One of them, called 'Macho B', had been previously photographed in 1996 in the area. For any permanent population in the USA to thrive, protection from killing, an adequate prey base, and connectivity with Mexican populations are essential. In February 2009, a 53.5 kg (118 lb) jaguar was caught, radio-collared and released in an area southwest of Tucson, Arizona; this is farther north than had previously been expected and represents a sign there may be a permanent breeding population of jaguars within southern Arizona. The animal was later confirmed to be indeed the same male individual ('Macho B') that was photographed in 2004. On 2 March 2009, Macho B was recaptured and euthanized after he was found to be suffering from kidney failure; the animal was thought to be 16 years old, older than any known wild jaguar.
Completion of the United States–Mexico barrier as currently proposed will reduce the viability of any population currently residing in the United States, by reducing gene flow with Mexican populations, and prevent any further northward expansion for the species.
The historic range of the species included much of the southern half of the United States, and in the south extended much farther to cover most of the South American continent. In total, its northern range has receded 1,000 km (621 mi) southward and its southern range 2,000 km (1243 mi) northward. Ice age fossils of the jaguar, dated between 40,000 and 11,500 years ago, have been discovered in the United States, including some at an important site as far north as Missouri. Fossil evidence shows jaguars of up to 190 kg (420 lb), much larger than the contemporary average for the animal.
The habitat of the cat includes the rain forests of South and Central America, open, seasonally flooded wetlands, and dry grassland terrain. Of these habitats, the jaguar much prefers dense forest; the cat has lost range most rapidly in regions of drier habitat, such as the Argentine pampas, the arid grasslands of Mexico, and the southwestern United States. The cat will range across tropical, subtropical, and dry deciduous forests (including, historically, oak forests in the United States). The jaguar prefers to live by rivers, swamps, and in dense rainforest with thick cover for stalking prey. Jaguars have been found at elevations as high as 3,800 m, but they typically avoid montane forest and are not found in the high plateau of central Mexico or in the Andes. The jaguars preferred habitats are usually swamps and wooded regions, but jaguars also live in scrublands and deserts.
Ecology and behavior
The adult jaguar is an apex predator, meaning it exists at the top of its food chain and is not preyed on in the wild. The jaguar has also been termed a keystone species, as it is assumed, through controlling the population levels of prey such as herbivorous and granivorous mammals, apex felids maintain the structural integrity of forest systems. However, accurately determining what effect species like the jaguar have on ecosystems is difficult, because data must be compared from regions where the species is absent as well as its current habitats, while controlling for the effects of human activity. It is accepted that mid-sized prey species undergo population increases in the absence of the keystone predators, and this has been hypothesized to have cascading negative effects. However, field work has shown this may be natural variability and the population increases may not be sustained. Thus, the keystone predator hypothesis is not accepted by all scientists.
The jaguar also has an effect on other predators. The jaguar and the cougar, the next-largest feline of the Americas, are often sympatric (related species sharing overlapping territory) and have often been studied in conjunction. Where sympatric with the jaguar, the cougar is smaller than normal and is smaller than the local jaguars. The jaguar tends to take larger prey, usually over 22 kg (49 lb) and the cougar smaller, usually between 2 and 22 kg (4.4 and 48.5 lb), reducing the latter's size. This situation may be advantageous to the cougar. Its broader prey niche, including its ability to take smaller prey, may give it an advantage over the jaguar in human-altered landscapes; while both are classified as near-threatened species, the cougar has a significantly larger current distribution.
Reproduction and life cycle
Jaguar females reach sexual maturity at about two years of age, and males at three or four. The cat is believed to mate throughout the year in the wild, although births may increase when prey is plentiful. Research on captive male jaguars supports the year-round mating hypothesis, with no seasonal variation in semen traits and ejaculatory quality; low reproductive success has also been observed in captivity. Female estrus is 6–17 days out of a full 37-day cycle, and females will advertise fertility with urinary scent marks and increased vocalization. Both sexes will range more widely than usual during courtship.
Pairs separate after mating, and females provide all parenting. The gestation period lasts 93–105 days; females give birth to up to four cubs, and most commonly to two. The mother will not tolerate the presence of males after the birth of cubs, given a risk of infanticide; this behavior is also found in the tiger.
The young are born blind, gaining sight after two weeks. Cubs are weaned at three months, but remain in the birth den for six months before leaving to accompany their mother on hunts. They will continue in their mother's company for one to two years before leaving to establish a territory for themselves. Young males are at first nomadic, jostling with their older counterparts until they succeed in claiming a territory. Typical lifespan in the wild is estimated at around 12–15 years; in captivity, the jaguar lives up to 23 years, placing it among the longest-lived cats.
Like most cats, the jaguar is solitary outside mother–cub groups. Adults generally meet only to court and mate (though limited noncourting socialization has been observed anecdotally) and carve out large territories for themselves. Female territories, which range from 25 to 40 km2 in size, may overlap, but the animals generally avoid one another. Male ranges cover roughly twice as much area, varying in size with the availability of game and space, and do not overlap. The territory of a male can contain those of several females. The jaguar uses scrape marks, urine, and feces to mark its territory.
Like the other big cats, the jaguar is capable of roaring and does so to warn territorial and mating competitors away; intensive bouts of counter-calling between individuals have been observed in the wild. Their roar often resembles a repetitive cough, and they may also vocalize mews and grunts. Mating fights between males occur, but are rare, and aggression avoidance behavior has been observed in the wild. When it occurs, conflict is typically over territory: a male's range may encompass that of two or three females, and he will not tolerate intrusions by other adult males.
The jaguar is often described as nocturnal, but is more specifically crepuscular (peak activity around dawn and dusk). Both sexes hunt, but males travel farther each day than females, befitting their larger territories. The jaguar may hunt during the day if game is available and is a relatively energetic feline, spending as much as 50–60 percent of its time active. The jaguar's elusive nature and the inaccessibility of much of its preferred habitat make it a difficult animal to sight, let alone study.
Hunting and diet
Like all cats, the jaguar is an obligate carnivore, feeding only on meat. It is an opportunistic hunter and its diet encompasses at least 87 species. The jaguar can take virtually any terrestrial or riparian vertebrate found in Central or South America, with a preference for large prey. The jaguar is more of a dietary generalist than its Old World cousins: the American tropics have a high diversity of small animals but relatively low populations and diversity of the large ungulates which this genus favors. They regularly take adult caimans, deer, capybaras, tapirs, peccaries, dogs, zorros, and sometimes even anacondas. However, the cat will eat any small species that can be caught, including frogs, mice, birds (mainly ground-based species such as cracids), fish, sloths, monkeys, and turtles; a study conducted in Cockscomb Basin Wildlife Sanctuary in Belize, for example, revealed that the diets of jaguars there consisted primarily of armadillos and pacas. Some jaguars will also take domestic livestock. El Jefe, one of the few jaguars that were reported in the United States, has also been found to kill and eat American black bears, as deduced from hairs found within his scats and the partly consumed carcass of a black bear sow with the distinctive puncture marks to the skull left by jaguars. This indicates that jaguars might have once preyed on black bears when the species was still present in the area. Spectacled bears are also known to avoid jaguars, possibly because they may constitute occasional prey items.
While the jaguar often employs the deep throat-bite and suffocation technique typical among Panthera, it sometimes uses a killing method unique amongst cats: it pierces directly through the temporal bones of the skull between the ears of prey (especially the capybara) with its canine teeth, piercing the brain. This may be an adaptation to "cracking open" turtle shells; following the late Pleistocene extinctions, armored reptiles such as turtles would have formed an abundant prey base for the jaguar. The skull bite is employed with mammals in particular; with reptiles such as the caiman, the jaguar may leap onto the back of the prey and sever the cervical vertebrae, immobilizing the target. When attacking sea turtles, including the huge leatherback sea turtle which weighs about 385 kg (849 lb) on average, as they try to nest on beaches, the jaguar will bite at the head, often beheading the prey, before dragging it off to eat. Reportedly, while hunting horses, a jaguar may leap onto their back, place one paw on the muzzle and another on the nape and then twist, dislocating the neck. Local people have anecdotally reported that when hunting a pair of horses bound together, the jaguar will kill one horse and then drag it while the other horse, still living, is dragged in their wake. With prey such as smaller dogs, a paw swipe to the skull may be sufficient to kill it.
The jaguar is a stalk-and-ambush rather than a chase predator. The cat will walk slowly down forest paths, listening for and stalking prey before rushing or ambushing. The jaguar attacks from cover and usually from a target's blind spot with a quick pounce; the species' ambushing abilities are considered nearly peerless in the animal kingdom by both indigenous people and field researchers, and are probably a product of its role as an apex predator in several different environments. The ambush may include leaping into water after prey, as a jaguar is quite capable of carrying a large kill while swimming; its strength is such that carcasses as large as a heifer can be hauled up a tree to avoid flood levels.
On killing prey, the jaguar will drag the carcass to a thicket or other secluded spot. It begins eating at the neck and chest, rather than the midsection. The heart and lungs are consumed, followed by the shoulders. The daily food requirement of a 34 kg (75 lb) animal, at the extreme low end of the species' weight range, has been estimated at 1.4 kg (3.1 lb). For captive animals in the 50–60 kg (110–130 lb) range, more than 2 kg (4.4 lb) of meat daily are recommended. In the wild, consumption is naturally more erratic; wild cats expend considerable energy in the capture and kill of prey, and they may consume up to 25 kg (55 lb) of meat at one feeding, followed by periods of famine.
Unlike all other Panthera species, jaguars very rarely attack humans. However, jaguar attacks appear to be on the rise with increased human encroachment on their habitat and a decrease in prey populations. Sometimes jaguars in captivity attack zookeepers. In addition, it appears that attacks on humans had been more common in the past, at least after conquistadors arrived in the Americas, to the extent that the jaguar had a fearsome reputation in the Americas, akin to the lion and tiger in the Old World. Nevertheless, even in those times, the jaguar's chief prey was the capybara, not the human, and Charles Darwin reported a saying of Native Americans that people would not have to fear the jaguar much, as long as capybaras were abundant.
Jaguar populations are rapidly declining. The animal is considered Near Threatened by the International Union for Conservation of Nature (IUCN), meaning it may be threatened with extinction in the near future. The loss of parts of its range, including its virtual elimination from its historic northern areas and the increasing fragmentation of the remaining range, have contributed to this status. The 1960s had particularly significant declines, with more than 15,000 jaguar skins brought out of the Brazilian Amazon yearly; the Convention on International Trade in Endangered Species of 1973 brought about a sharp decline in the pelt trade. Detailed work performed under the auspices of the Wildlife Conservation Society revealed the specieshas lost 37% of its historic range, with its status unknown in an additional 18 percent. More encouragingly, the probability of long-term survival was considered high in 70% of its remaining range, particularly in the Amazon basin and the adjoining Gran Chaco and Pantanal.
The major risks to the jaguar include deforestation across its habitat, increasing competition for food with human beings, especially in dry and unproductive habitat, poaching, hurricanes in northern parts of its range, and the behavior of ranchers who will often kill the cat where it preys on livestock. When adapted to the prey, the jaguar has been shown to take cattle as a large portion of its diet; while land clearance for grazing is a problem for the species, the jaguar population may have increased when cattle were first introduced to South America, as the animals took advantage of the new prey base. This willingness to take livestock has induced ranch owners to hire full-time jaguar hunters.
The skins of wild cats and other mammals have been highly valued by the fur trade for many decades. From the beginning of the 20th-century Jaguars were hunted in large numbers, but over-harvest and habitat destruction reduced the availability and induced hunters and traders to gradually shift to smaller species by the 1960s. The international trade of jaguar skins had its largest boom between the end of the Second World War and the early 1970, due to the growing economy and lack of regulations. From 1967 onwards, the regulations introduced by national laws and international agreements diminished the reported international trade from as high as 13000 skins in 1967, through 7000 skins in 1969, until it became negligible after 1976, although illegal trade and smuggling continue to be a problem. During this period, the biggest exporters were Brazil and Paraguay, and the biggest importers were the USA and Germany.
The jaguar is listed on CITES Appendix I, which means that all international trade in jaguars or their parts is prohibited. Hunting jaguars is prohibited in Argentina, Brazil, Colombia, French Guiana, Honduras, Nicaragua, Panama, Paraguay, Suriname, the United States, and Venezuela. Hunting of jaguars is restricted in Brazil, Costa Rica, Guatemala, Mexico and Peru. Trophy hunting is still permitted in Bolivia, and it is not protected in Ecuador or Guyana. In the US, it is listed as endangered under the Endangered Species Act.
Jaguar Conservation Units
Jaguar conservation is complicated because of the species' large range spaning 18 countries with different policies and regulations. Specific areas of high importance for jaguar conservation, so-called "Jaguar Conservation Units" (JCU) were determined in 2000. These are large areas inhabited by at least 50 jaguars. Each unit was assessed and evaluated on the basis of size, connectivity, habitat quality for both jaguar and prey, and jaguar population status. That way, 51 Jaguar Conservation Units were determined in 36 geographic regions as priority areas for jaguar conservation including:
- the Sierra Madre of Mexico
- the Selva Maya tropical forests extending over Mexico, Belize, and Guatemala
- the Chocó-Darién moist forests from Honduras, Panama to Colombia
- Sierra de Tamaulipas
- Venezuelan Llanos
- northern Cerrado and Amazon basin in Brazil
- Misiones Province in Argentina
Recent studies underlined that to maintain the robust exchange across the jaguar gene pool necessary for maintaining the species, it is important that jaguar habitats are interconnected. To facilitate this, a new project, the Paseo del Jaguar, has been established to connect several jaguar hotspots.
Given the inaccessibility of much of the species' range, particularly the central Amazon, estimating jaguar numbers is difficult. Researchers typically focus on particular bioregions, thus species-wide analysis is scant. In 1991, 600–1,000 (the highest total) were estimated to be living in Belize. A year earlier, 125–180 jaguars were estimated to be living in Mexico's 4,000-km2 (2400-mi2) Calakmul Biosphere Reserve, with another 350 in the state of Chiapas. The adjoining Maya Biosphere Reserve in Guatemala, with an area measuring 15,000 km2 (9,000 mi2), may have 465–550 animals. Work employing GPS telemetry in 2003 and 2004 found densities of only six to seven jaguars per 100 km2 in the critical Pantanal region, compared with 10 to 11 using traditional methods; this suggests the widely used sampling methods may inflate the actual numbers of cats.
In setting up protected reserves, efforts generally also have to be focused on the surrounding areas, as jaguars are unlikely to confine themselves to the bounds of a reservation, especially if the population is increasing in size. Human attitudes in the areas surrounding reserves and laws and regulations to prevent poaching are essential to make conservation areas effective.
To estimate population sizes within specific areas and to keep track of individual jaguars, camera trapping and wildlife tracking telemetry are widely used, and feces may be sought out with the help of detector dogs to study jaguar health and diet. Current conservation efforts often focus on educating ranch owners and promoting ecotourism. The jaguar is generally defined as an umbrella species – its home range and habitat requirements are sufficiently broad that, if protected, numerous other species of smaller range will also be protected. Umbrella species serve as "mobile links" at the landscape scale, in the jaguar's case through predation. Conservation organizations may thus focus on providing viable, connected habitat for the jaguar, with the knowledge other species will also benefit.
Ecotourism setups are being used to generate public interest in charismatic animals such as the jaguar, while at the same time generating revenue that can be used in conservation efforts. Audits done in Africa have shown that ecotourism has helped in African cat conservation. As with large African cats, a key concern in jaguar ecotourism is the considerable habitat space the species requires, so if ecotourism is used to aid in jaguar conservation, some considerations need to be made as to how existing ecosystems will be kept intact, or how new ecosystems that are large enough to support a growing jaguar population will be put into place.
The United States
The only extant cat native to North America that roars, the jaguar was recorded as an animal of the Americas by Thomas Jefferson in 1799. Jaguars are still occasionally sighted in Arizona, such as El Jefe, prompting actions for its conservation by authorities. For example, on the 20th of August, 2012, the USFWS proposed setting aside 838,232 acres in Arizona and New Mexico — an area larger than Rhode Island — as critical jaguar habitat.
In mythology and culture
In pre-Columbian Central and South America, the jaguar was a symbol of power and strength. Among the Andean cultures, a jaguar cult disseminated by the early Chavín culture became accepted over most of what is today Peru by 900 BC. The later Moche culture of northern Peru used the jaguar as a symbol of power in many of their ceramics. In the religion of the Muisca, who inhabited the cool Altiplano Cundiboyacense in the Colombian Andes, the jaguar was considered a sacred animal and during their religious rituals the people dressed in jaguar skins. The skins were traded with the lowland peoples of the tropical Llanos Orientales. The name of zipa Nemequene was derived from the Muysccubun words nymy and quyne, meaning "force of the jaguar".
In Mesoamerica, the Olmec—an early and influential culture of the Gulf Coast region roughly contemporaneous with the Chavín—developed a distinct "were-jaguar" motif of sculptures and figurines showing stylised jaguars or humans with jaguar characteristics. In the later Maya civilization, the jaguar was believed to facilitate communication between the living and the dead and to protect the royal household. The Maya saw these powerful felines as their companions in the spiritual world, and a number of Maya rulers bore names that incorporated the Mayan word for jaguar (b'alam in many of the Mayan languages). Balam (Jaguar) remains a common Maya surname, and it is also the name of Chilam Balam, a legendary author to whom are attributed 17th and 18th-centuries Maya miscellanies preserving much important knowledge. The Aztec civilization shared this image of the jaguar as the representative of the ruler and as a warrior. The Aztecs formed an elite warrior class known as the Jaguar Knights. In Aztec mythology, the jaguar was considered to be the totem animal of the powerful deity Tezcatlipoca.
The jaguar and its name are widely used as a symbol in contemporary culture. It is the national animal of Guyana, and is featured in its coat of arms. The flag of the Department of Amazonas, a Colombian department, features a black jaguar silhouette pouncing towards a hunter. The jaguar also appears in banknotes of Brazilian real. The jaguar is also a common fixture in the mythology of many contemporary native cultures in South America, usually being portrayed as the creature which gave humans the power over fire.
Jaguar is widely used as a product name, most prominently for a British luxury car brand. The name has been adopted by sports franchises, including the NFL's Jacksonville Jaguars and the Mexican soccer club Chiapas F.C. The crest of Argentina's national federation in rugby union features a jaguar; however, because of a journalist error, the country's national team is nicknamed Los Pumas. In the spirit of the ancient Mayan culture, the 1968 Olympics in Mexico City adopted a red jaguar as the first official Olympic mascot.
- Caso, A.; Lopez-Gonzalez, C.; Payan, E.; Eizirik, E.; de Oliveira, T.; Leite-Pitman, R.; Kelly, M. & Valderrama, C. (2008). "Panthera onca". IUCN Red List of Threatened Species. Version 2017-2. International Union for Conservation of Nature.
- Wroe, S.; McHenry, C.; Thomason, J. (2006). "Bite club: comparative bite force in big biting mammals and the prediction of predatory behavior in fossil taxa" (PDF). Proceedings of the Royal Society B. 272 (1563): 619–25. doi:10.1098/rspb.2004.2986. PMC . PMID 15817436. Archived from the original (PDF) on 21 September 2006.
- Emmons, L.H. (1987). "Comparative feeding ecology of felids in a neotropical rainforest". Behavioral Ecology and Sociobiology 20 (4): 271–283.
- Rosa, C. L. de la & Nocke, C. C. (2000). "Jaguar (Panthera onca)". A guide to the carnivores of Central America: natural history, ecology, and conservation. University of Texas Press. pp. 39–?. ISBN 978-0-292-71604-9.
- Harper, D. (2001–2017). "Jaguar". Online Etymology Dictionary. Douglas Harper.
- "Breve Vocabulario" (in Spanish). Faculty of Law, University of Buenos Aires. Archived from the original on 2006-09-01. Retrieved 2006-09-29.
- Díaz, Eduardo Acevedo (1890). "Notas". Nativas (in Spanish). Retrieved 2006-09-29.
- "Yaguareté – La Verdadera Fiera". RED Yaguareté (in Spanish). Archived from the original on 2006-09-03. Retrieved 2006-09-27.
- Liddell, H. G. & R. Scott (1940). "πάνθηρ". A Greek-English Lexicon, revised and augmented. Oxford: Clarendon Press.
- OUP (2002). "Panther". Oxford English Dictionary. Oxford: Oxford University Press.
- "The Jaguar in Mexico". Mexicolore.
- "ounce" 2, Oxford English Dictionary, 2nd edition
- Turner, A. (1987). "New fossil carnivore remains from the Sterkfontein hominid site (Mammalia: Carnivora)". Annals of the Transvaal Museum. 34: 319–347. ISSN 0041-1752.
- Johnson, W. E.; Eizirik, E.; Pecon-Slattery, J.; Murphy, W. J.; Antunes, A.; Teeling, E.; O'Brien, S. J. (2006). "The Late Miocene radiation of modern Felidae: A genetic assessment". Science. 311 (5757): 73–77. Bibcode:2006Sci...311...73J. doi:10.1126/science.1122277. PMID 16400146.
- Eizirik, E.; Kim, J. H.; Menotti-Raymond, M.; Crawshaw P. G. Jr.; O'Brien, S. J.; Johnson, W. E. (2001). "Phylogeography, population history and conservation genetics of jaguars (Panthera onca, Mammalia, Felidae)". Molecular Ecology. 10 (1): 65–79. doi:10.1046/j.1365-294X.2001.01144.x. PMID 11251788.
- Janczewski, D. N.; Modi, W. S.; Stephens, J. C.; O'Brien, S. J. (1996). "Molecular evolution of mitochondrial 12S RNA and cytochrome b sequences in the pantherine lineage of Felidae". Molecular Biology and Evolution. 12 (4): 690–707. doi:10.1093/oxfordjournals.molbev.a040232. PMID 7544865.
- Johnson, W. E.; O'Brien, S. J. (1997). "Phylogenetic reconstruction of the Felidae using 16S rRNA and NADH-5 mitochondrial genes". Journal of Molecular Evolution. 44: S98–116. doi:10.1007/PL00000060. PMID 9071018.
- Yu, L.; Zhang, Y. P. (2005). "Phylogenetic studies of pantherine cats (Felidae) based on multiple genes, with novel application of nuclear beta-fibrinogen intron 7 to carnivores". Molecular Phylogenetics and Evolution. 35 (2): 483–495. doi:10.1016/j.ympev.2005.01.017. PMID 15804417.
- Wozencraft, W.C. (2005). "Order Carnivora". In Wilson, D.E.; Reeder, D.M. Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 546–547. ISBN 978-0-8018-8221-0. OCLC 62265494.
- Pocock, R. I. (1939). "The races of jaguar (Panthera onca)". Novitates Zoologicae. 41: 406–422.
- Seymour, K.L. (1989). "Panthera onca" (PDF). Mammalian Species. 340 (340): 1–9. doi:10.2307/3504096. JSTOR 3504096. Retrieved 2009-12-27.
- Larson, S. E. (1997). "Taxonomic re-evaluation of the jaguar". Zoo Biology. 16 (2): 107–120. doi:10.1002/(SICI)1098-2361(1997)16:2<107::AID-ZOO2>3.0.CO;2-E.
- Ruiz-Garcia, M.; Payan, E; Murillo, A.; Alvarez, D. (2006). "DNA microsatellite characterization of the jaguar (Panthera onca) in Colombia" (PDF). Genes & Genetic Systems. 81 (2): 115–127. doi:10.1266/ggs.81.115. Retrieved 2011-11-11.
- Kitchener, A. C., Breitenmoser-Würsten, C., Eizirik, E., Gentry, A., Werdelin, L., Wilting A., Yamaguchi, N., Abramov, A. V., Christiansen, P., Driscoll, C., Duckworth, J. W., Johnson, W., Luo, S.-J., Meijaard, E., O’Donoghue, P., Sanderson, J., Seymour, K., Bruford, M., Groves, C., Hoffmann, M., Nowell, K., Timmons, Z. & Tobe, S. (2017). "A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group" (PDF). Cat News. Special Issue 11.
- Anton, Mauricio (1997). The Big Cats and Their Fossil Relatives. Columbia University Press. p. 63. ISBN 978-0-231-10228-5.
- Nowak, Ronald M (1999). Walker's Mammals of the World. 2. Baltimore: Johns Hopkins University Press. p. 831. ISBN 0-8018-5789-9.
- Burnien, David; Wilson, Don E. (2001). Animal: The Definitive Visual Guide to the World's Wildlife. New York City: Dorling Kindersley. ISBN 0-7894-7764-5.
- "The Jaguar (Panthera onca) – Information and Data".
- Boitani, Luigi (1984). Simon and Schuster's Guide to Mammals. Simon & Schuster. ISBN 0-671-43727-5.
- "All about Jaguars: Ecology". Wildlife Conservation Society. Archived from the original on 2009-05-29. Retrieved 2006-08-11.
- Rodrigo Nuanaez; Brian Miller; Fred Lindzey (2000). "Food habits of jaguars and pumas in Jalisco, Mexico". Journal of Zoology. 252 (3): 373–379. doi:10.1111/j.1469-7998.2000.tb00632.x. Retrieved 2006-08-08.
- "Jaguar Fact Sheet". Jaguar Species Survival Plan. American Zoo and Aquarium Association. Archived from the original on 2012-01-27. Retrieved 2006-08-14.
- King Of Cats (21 January 2010). "142 kg Wild Jaguar" – via YouTube.
- Nowell, K.; Jackson, P., eds. (1996). "Panthera Onca". Wild Cats. Status Survey and Conservation Action Plan (PDF). Gland, Switzerland: IUCN/SSC Cat Specialist Group. IUCN. pp. 118–122. Retrieved 2011-11-11.
- Hartstone-Rose, Adam; Perry, Jonathan M. G.; Morrow, Caroline J. (1 August 2012). "Bite Force Estimation and the Fiber Architecture of Felid Masticatory Muscles". Anat Rec. 295 (8): 1336–1351. doi:10.1002/ar.22518 – via Wiley Online Library.
- "Search for the Jaguar". National Geographic Specials. Kentucky Educational Television. 2003. Archived from the original on 20 July 2013. Retrieved 19 March 2012.
- McGrath, Susan (August 2004). "Top Cat". National Audubon Society. Archived from the original on 4 July 2008. Retrieved 2009-12-02.
- "Jaguar (Panthera onca)". Our animals. Akron Zoo. Archived from the original on 2011-07-24. Retrieved 2006-08-11.
- Dinets, Vladmir. "First documentation of melanism in the jaguar (Panthera onca) from northern Mexico". Archived from the original on 26 September 2006. Retrieved 29 September 2006.
- Meyer, John R. (1994). "Black jaguars in Belize?: A survey of melanism in the jaguar, Panthera onca". Belize Explorer Group. biological-diversity.info.
- U.S. Fish and Wildlife Service, 2016, Jaguar Draft Recovery Plan, p.5
- Brulliard, Karen (December 8, 2016). "Jaguar spotting: A new wild cat may be roaming the United States". The Washington Post. Retrieved May 21, 2017.
- "Determination That Designation of Critical Habitat Is Not Prudent for the Jaguar". Federal Register Environmental Documents. 2006-07-12. Retrieved 2006-08-30.
- Dye, Lee (February 21, 2002). "Jaguars Moving Back Into U.S." abcnews.go.com. ABC News. Retrieved May 21, 2017.
- Mccain, Emil B.; Childs, Jack L. (2008). "Evidence of resident Jaguars (Panthera onca) in the Southwestern United States and the Implications for Conservation" (PDF). Journal of Mammalogy. 89 (1): 1–10. doi:10.1644/07-MAMM-F-268.1. Archived from the original (PDF) on 18 January 2015. Retrieved 2011-11-11.
- "Jaguar Management". Arizona Game and Fish Department. 2009. Archived from the original on 4 October 2006. Retrieved 8 August 2006.
- "Arizona Game and Fish collars first wild jaguar in United States". Readitnews.com. Retrieved 2009-03-08.
- Hock, Heather (2009-03-02). "Illness forced vets to euthuanize recaptured jaguar". Azcentral.com. Retrieved 2009-03-08.
- "Addressing the Impacts of Border Security Activities On Wildlife and Habitat in Southern Arizona: Stakeholder Recommendations" (PDF). Wildlands Project. Archived from the original (PDF) on 2007-07-11. Retrieved 2008-11-03.
- "Jaguars". The Midwestern United States 16,000 years ago. Illinois State Museum. Retrieved 2006-08-20.
- "Jaguar". Encyclopædia Britannica.
- "Jaguar (Panthera Onca)" (PDF). Phoenix Zoo. Archived from the original (PDF) on 15 December 2014. Retrieved 30 August 2006.
- "Structure and Character: Keystone Species". Mongabay. Archived from the original on 2014-10-06. Retrieved 2006-08-30.
- Wright, S. J.; Gompper, M. E.; DeLeon, B. (1994). "Are large predators keystone species in Neotropical forests? The evidence from Barro Colorado Island". Oikos. 71 (2): 279–294. doi:10.2307/3546277. JSTOR 3546277. Archived from the original on 12 October 2007. Retrieved 11 November 2011.
- Brakefield, T. (1993). Big Cats: Kingdom of Might. ISBN 0-89658-329-5.
- Iriarte, J. A.; Franklin, W. L.; Johnson, W. E.; Redford, K. H. (1990). "Biogeographic variation of food habits and body size of the America puma". Oecologia. 85 (2): 185–190. doi:10.1007/BF00319400.
- Baker, Reproduction, pp. 28–38.
- Morato, R. G.; Vaz Guimaraes, M; A; B.; Ferriera, F.; Nascimento Verreschi, I. T.; Renato Campanarut Barnabe (1999). "Reproductive characteristics of captive male jaguars". Brazilian Journal of Veterinary Research and Animal Science. 36 (5). Retrieved 2011-11-11.
- Baker, Natural History and Behavior, pp. 8–16.
- Jeff Egerton (Spring 2006). "Jaguars: Magnificence in the Southwest" (PDF). Newsletter. Southwest Wildlife Rehabilitation & Educational Foundation. Archived from the original (PDF) on 21 July 2011. Retrieved 2009-12-06.
- Schaller, George B.; Crawshaw, Peter Gransden Jr. (1980). "Movement Patterns of Jaguar". Biotropica. 12 (3): 161–168. doi:10.2307/2387967. JSTOR 2387967.
- Rabinowitz, A. R.; Nottingham, B. G. Jr (1986). "Ecology and behaviour of the Jaguar (Panthera onca) in Belize, Central America". Journal of Zoology. 210 (1): 149–159. doi:10.1111/j.1469-7998.1986.tb03627.x. Overlapping male ranges are observed in this study in Belize. Note the overall size of ranges is about half of normal.
- Harmsen, Bart J.; Foster, Rebecca J.; Gutierrez, Said M.; Marin, Silverio Y.; Doncaster, C. Patrick. "Scrape-marking behavior of jaguars (Panthera onca) and pumas (Puma concolor)". Journal of Mammalogy. 91 (5): 1225–1234. doi:10.1644/09-mamm-a-416.1.
- Weissengruber, G. E.; Forstenpointner, G.; Peters, G.; Kübber-Heiss, A.; Fitch, W. T. (2002). "Hyoid apparatus and pharynx in the lion (Panthera leo), jaguar (Panthera onca), tiger (Panthera tigris), cheetah (Acinonyx jubatus) and domestic cat (Felis silvestris f. catus)". Journal of Anatomy. 201 (3): 195–209. doi:10.1046/j.1469-7580.2002.00088.x. PMC . PMID 12363272.
- Hast, M. H. (1989). "The larynx of roaring and non-roaring cats". Journal of Anatomy. 163: 117–121. PMC . PMID 2606766.
- Otfinoski, Steven (2010). Jaguars. Marshall Cavendish. p. 18. ISBN 978-0-7614-4839-6. Retrieved 2011-03-16.
- "Jaguar". Kids' Planet. Defenders of Wildlife. Archived from the original on 30 September 2006. Retrieved 23 September 2006.
- "Rare jaguar sighting in Arizona, 60 miles north of Mexican border". Newsweek. 2017. Retrieved 2017-03-03.
- Richard Grant (October 2016). "The Return of the Great American Jaguar". Smithsonian Magazine. Retrieved 2016-10-02.
- Servheen, C.; Herrero, S.; Peyton, B. (1999). Bears: Status Survey and Conservation Action Plan (PDF). Missoula, Montana: IUCN/SSC Bear Specialist Group. ISBN 978-2-8317-0462-3.
- Robin Rood (2011). "Reassessing the Cultural and Psychopharmacological Significance of Banisteriopsis caapi: Preparation, Classification and Use Among the Piaroa of Southern Venezuela". Journal of Psychoactive Drugs. 40 (3): 301–307. doi:10.1080/02791072.2008.10400645.
- Schaller, G. B.; Vasconselos, J. M. C. (1978). "Jaguar predation on capybara" (PDF). Zeitschrift für Säugetierkunde. 43: 296–301. Archived from the original (PDF) on 31 March 2012. Retrieved 2011-11-11.
- Travellers' Wildlife Guide to Costa Rica by Les Beletsky. Interlink Publishing Group (2004), ISBN 1566565294
- The animal kingdom: based upon the writings of the eminent naturalists Audubon, Wallace, Brehm, Wood, and Others, edited by Hugh Craig. Trinity College (1897), New York.
- Baker, Hand-rearing, pp. 62–75 (table 5).
- Baker, Nutrition, pp. 55–61.
- V. Iserson, K.; Francis, Adama M. (2015). "Jaguar Attack on a Child: Case Report and Literature Review". Western Journal of Emergency Medicine. 16 (2): 303–309. doi:10.5811/westjem.2015.1.24043. PMC . PMID 25834674.
- "Jaguar: The Western Hemisphere's Top Cat". Planeta. February 2008. Archived from the original on 21 August 2008. Retrieved 8 March 2009.
- Porter, J. H. (1894). "The Jaguar". Wild beasts; a study of the characters and habits of the elephant, lion, leopard, panther, jaguar, tiger, puma, wolf, and grizzly bear. pp. 174–195. Retrieved 2014-01-19.
- Weber, William; Rabinowitz, Alan (August 1996). "A Global Perspective on Large Carnivore Conservation" (PDF). Conservation Biology. 10 (4): 1046–1054. doi:10.1046/j.1523-1739.1996.10041046.x. Archived from the original (PDF) on 26 April 2012. Retrieved 17 December 2009.
- Sanderson, E. W.; K. H., Redford; Chetkiewicz, C. B.; Medellin, R. A.; Rabinowitz, A. R.; Robinson, J. G.; Taber, A. B. (2002). "Planning to save a species: the jaguar as a model". Conservation Biology. 16 (1): 58–72. doi:10.1046/j.1523-1739.2002.00352.x.
- Jędrzejewski, Włodzimierz; Boede, Ernesto O.; Abarca, María; Sánchez-Mercado, Ada; Ferrer-Paris, José R.; Lampo, Margarita; Velásquez, Grisel; Carreño, Rafael; Viloria, Ángel L.; Hoogesteijn, Rafael; Robinson, Hugh S.; Stachowicz, Izabela; Cerda, Hugo; Weisz, María del Mar; Barros, Tito R.; Rivas, Gilson A.; Borges, Gilberto; Molinari, Jesús; Lew, Daniel; Takiff, Howard; Schmidt, Krzysztof (2017). "Predicting carnivore distribution and extirpation rate based on human impacts and productivity factors; assessment of the state of jaguar (Panthera onca) in Venezuela". Biological Conservation. 206: 132–142. doi:10.1016/j.biocon.2016.09.027. Retrieved 5 January 2017.
- Broad, Steven (1987). The harvest of and trade in Latin American spotted cats (Felidae) and otters (Lutrinae). Cambridge, UK: IUCN Conservation Monitoring Centre. Retrieved 8 July 2016.
- "Path of the jaguars project". Ngm.nationalgeographic.com. March 2009. Retrieved 2010-04-02.
- Weckel, M., Giuliano, W. and Silver, S. (2006). "Cockscomb revisited: jaguar diet in the Cockscomb Basin Wildlife Sanctuary, Belize". Biotropica 38 (5): 687–690.
- Baker, Protection and Population Status, p. 4.
- Soisalo, M. K.; Cavalcanti, S. M. C. (2006). "jaguar "Estimating the density of a jaguar population in the Brazilian Pantanal using camera-traps and capture–recapture sampling in combination with GPS radio-telemetry" (PDF). Biological Conservation. 129 (4): 487–496. doi:10.1016/j.biocon.2005.11.023. Archived from the original (PDF) on 21 September 2006. Retrieved 2006-08-08.
- Gutierrez-Gonzalez, Carmina E.; Gomez-Ramirez, Miguel A.; Lopez-Gonzalez, Carlos A.; Doherty, Paul F. (2015). "Are Private Reserves Effective for Jaguar Conservation?". PLoS ONE. 10 (9): e0137541. doi:10.1371/journal.pone.0137541. PMC . PMID 26398115.
- Soisalo, Marianne K.; Cavalcanti, Sandra M.C. (2006). "Estimating the Density of a Jaguar Population in the Brazilian Pantanal Using Camera-traps and Capture-recapture Sampling in Combination with GPS Radio-telemetry". Biological Conservation. 129 (4): 487. doi:10.1016/j.biocon.2005.11.023.
- Furtado, M. M.; Carrillo-Percastegui, S. E.; Jácomo, A. T. A.; Powell, G.; Silveira, L.; Vynne, C.; Sollmann, R. (2008). "Studying jaguars in the wild: past experiences and future perspectives" (PDF). Cat News. 4 (special).
- "Jaguar Refuge in the Llanos Ecoregion". World Wildlife Fund. Archived from the original on 17 December 2014. Retrieved 1 September 2006.
- "Glossary". Sonoran Desert Conservation Plan: Kids. Pima County Government. Archived from the original on 2006-06-20. Retrieved 2006-09-01.
- Mossaz, A.; Buckley, R.C.; Castley, J.G. (2015). "Ecotourism Contributions to Conservation of African Big Cats". Journal for Nature Conservation. 28: 112. doi:10.1016/j.jnc.2015.09.009.
- Christie, Bob (2011-12-01). "Excitement follows 2 jaguar sighting in Arizona". The Arizona Republic. Archived from the original on 7 March 2015.
- "Full text of "The writings of Thomas Jefferson"".
- Jessica Lamberton-Moreno (September 8, 2015). "Student project results in new jaguar sighting". Sky Island Alliance. Retrieved 2015-01-04.
- Handwerk, Brian (3 February 2016). "Only Known Jaguar in U.S. Filmed in Rare Video". National Geographic News. Retrieved 2016-02-04.
- Greenberg, Susan H. (2012-08-21). "Kitty Corner: Jaguars Win Critical Habitat in U.S." Scientific American. Archived from the original on 2014-10-07. Retrieved 2012-08-25.
- Museo Arqueologico Rafael Larco Herrera (1997). Berrin, Katherine, ed. The Spirit of Ancient Peru: Treasures from the Museo Arqueologico Rafael Larco Herrera. New York City: Thames and Hudson. ISBN 0-500-01802-2.
- Bulliet, Richard W.; et al. (2010). The Earth and Its Peoples: A Global History. Cengage Learning. pp. 75–. ISBN 978-1-4390-8476-2. Retrieved 11 December 2011.
- Lockard, Craig A. (2010). Societies, Networks, and Transitions, Volume I: To 1500: A Global History. Cengage Learning. pp. 215–. ISBN 978-1-4390-8535-6. Retrieved 11 December 2011.
- Ocampo López, Javier (2007). Grandes culturas indígenas de América – Great indigenous cultures of the Americas (in Spanish). Bogotá, Colombia: Plaza & Janes Editores Colombia S.A. p. 231. ISBN 978-958-14-0368-4.
- Kruschek, Michael H. (2003). The evolution of the Bogotá chiefdom: A household view (PhD) (PDF). University of Pittsburgh. p. 14. Retrieved 11 January 2017.
- "nymy" (in Spanish). Muysccubun Dictionary Online. Retrieved 11 January 2017.
- "quyne" (in Spanish). Muysccubun Dictionary Online. Retrieved 11 January 2017.
- Christenson, Allen J. (2007). Popol Vuh: The Sacred Book of the Maya. University of Oklahoma Press. p. 196. ISBN 978-0-8061-3839-8. Retrieved 11 December 2011.
- "Guyana". RBC Radio. Archived from the original on 6 October 2014. Retrieved 11 November 2011.
- Gutterman, D. (2008-07-26). "Amazonas Department (Colombia)". Fotw.net. Archived from the original on 2011-08-05. Retrieved 2010-04-02.
- Levi-Strauss, Claude (2004) . O Cru e o Cozido. São Paulo: Cosac & Naify. Retrieved 2011-11-11.
- Davies, Sean (26 July 2007). "Puma power: Argentinian rugby". bbc.co.uk. Retrieved 8 October 2007.
- Welch, Paula. Cute Little Creatures: Mascots Lend a Smile to the Games (PDF). la84foundation.org. Retrieved 2011-11-11.
|Wikimedia Commons has media related to Panthera onca.|
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- "Species portrait Jaguar". International Union for Conservation of Nature Species Survival Commission Cat Specialist Group. Archived from the original on 2015-03-07.
- Sound of a jaguar roar at Vivanatura.org
- "Jaguar (Panthera onca)". Arkive. Archived from the original on 2014-12-28.
- "Jaguar". BBC. 2014. Archived from the original on 2014-09-22.
- People and Jaguars a Guide for Coexistence
- Sky Island Alliance website
- Felidae Conservation Fund
- "Jaguar". Encyclopedia Americana. 1920.