Maranhão red-handed howler

The Maranhão red-handed howler (Alouatta ululata) is an endangered species of howler monkey endemic to forests (for example Babaçu forests) in the northeastern Brazilian states of Ceará, Maranhão and Piauí.[1] It was previously thought to be a subspecies of the red-handed howler, but unlike that species, the Maranhão red-handed howler is strongly sexually dichromatic.[2]

Maranhão red-handed howler
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Primates
Suborder: Haplorhini
Infraorder: Simiiformes
Family: Atelidae
Genus: Alouatta
A. ululata
Binomial name
Alouatta ululata
Elliot, 1912
Maranhao Red-handed Howler area.png
Maranhao Red-handed Howler range

Alouatta belzebul ululata


  1. ^ a b de Oliveira, M.M. & Kierulff, M.C.M. (2008). "Alouatta ululata". The IUCN Red List of Threatened Species. IUCN. 2008: e.T918A13094890. doi:10.2305/IUCN.UK.2008.RLTS.T918A13094890.en. Retrieved 26 December 2017.
  2. ^ Gregorin, R. (2006). (in Portuguese) Taxonomia e variação geográfica das espécies do gênero Alouatta Lacépède (Primates, Atelidae) no Brasil. Rev. Bras. Zool. 23(1).