The Geophilidae are a polyphyletic, cosmopolitan family of soil centipedes in the superfamily Geophiloidea[3] containing the mostly defunct clades Aphilodontidae,[4] Dignathodontidae,[5] Linotaeniidae,[6] Chilenophilinae,[7] and Macronicophilidae.[8] Species in this family are characterized by mandibles with a single pectinate lamella, slender antennae, sternal pores with variable arrangement, a generally slightly or moderately elongate head, frequently undivided coxosternite with two paramedian sclerotized lines, claws without rows of filament, and female gonopods usually being an undivided lamina.[9]

Geophilus flavus (Geophilidae), Arnhem, the Netherlands.jpg
Geophilus flavus, the Netherlands
Scientific classification

Leach, 1815 [1]


Geophilus sp. mother and brood.

Compared to most other families in the suborder Adesmata, this family features a modest number of leg-bearing segments and limited variation in this number within each species.[10] Two species in this family include centipedes with only 29 pairs of legs: Geophilus persephones (29 pairs in the only specimen, a male), and G. richardi (29 or 31 pairs in the males and 33 pairs in the females).[11][12] In the order Geophilomorpha, only two species include centipedes with fewer leg pairs, both of them in the family Schendylidae.[11]

This family contains these genera:[1]


  1. ^ a b "Geophilidae Leach, 1815". Integrated Taxonomic Information System. Retrieved January 2, 2022.
  2. ^ "Geophilidae".
  3. ^ Bonato, Lucio (2014). "Phylogeny of Geophilomorpha (Chilopoda) inferred from new morphological and molecular evidence". Cladistics. 30 (5): 485–507. doi:10.1111/cla.12060. PMID 34794246. S2CID 86204188. Retrieved 27 October 2021.
  4. ^ Calvanese, Victor C. (2019-11-15). "Revision of the Neotropical species of Aphilodontinae (Geophilomorpha, Geophilidae), with eight new species and a first phylogenetic analysis of the subfamily". Zootaxa. 4698 (1): zootaxa.4698.1.1. doi:10.11646/zootaxa.4698.1.1. PMID 32229999. S2CID 209592191. Retrieved 21 December 2021.
  5. ^ "ITIS - Report: Dignathodontidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
  6. ^ "ITIS - Report: Linotaeniidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
  7. ^ Crabill, Ralph Edwin (1954). "A conspectus of the northeastern North American species of Geophilus (Chilopoda Geophilomorpha Geophilidae)". Proceedings of the Entomological Society of Washington. 56: 172–188. Retrieved 23 October 2021.
  8. ^ "ITIS - Report: Macronicophilidae". Integrated Taxonomic Information System. Retrieved 22 December 2021.
  9. ^ Minelli, Alessandro (2011-03-21). Treatise on Zoology - Anatomy, Taxonomy, Biology. The Myriapoda. Brill. p. 414. ISBN 978-90-04-18826-6. Retrieved 24 October 2021.
  10. ^ Minelli, Alessandro; Bortoletto, Stefano (1988-04-01). "Myriapod metamerism and arthropod segmentation". Biological Journal of the Linnean Society. 33 (4): 323–343. doi:10.1111/j.1095-8312.1988.tb00448.x. ISSN 0024-4066.
  11. ^ a b Pereira, Luis Alberto (2013-01-01). "Discovery of a second geophilomorph species (Myriapoda: Chilopoda) having twenty-seven leg-bearing segments, the lowest number recorded up to the present in the centipede order Geophilomorpha". Papéis Avulsos de Zoologia. 53 (13): 163–185. doi:10.1590/S0031-10492013001300001. ISSN 1807-0205.
  12. ^ Bonato, Lucio; Minelli, Alessandro; Drago, Leandro; Pereira, Luis Alberto (2015-09-25). "The phylogenetic position of Dinogeophilus and a new evolutionary framework for the smallest epimorphic centipedes (Chilopoda: Epimorpha)". Contributions to Zoology. 84 (3): 237–253. doi:10.1163/18759866-08403004. ISSN 1875-9866.