Amphilestes is a genus of extinct eutriconodont mammal from the Middle Jurassic of the United Kingdom. It was one of the first Mesozoic mammals discovered and described.

Temporal range: Bathonian
~168–166 Ma
Scientific classification edit
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Eutriconodonta
Family: Amphilestidae
Genus: Amphilestes
Owen, 1871
A. broderipii
Binomial name
Amphilestes broderipii
(Owen, 1871)


The first specimen of Amphilestes was discovered along with several other mammal jaws in the Stonesfield Slate Quarry, Oxfordshire before 1764.[1] However, it was not until 1812 that William Broderip bought the jaws, and he and his mentor - the famous palaeontologist Revd William Buckland - recognised that they were of mammal origin. Amphilestes broderipii was originally Amphitherium broderipii, until it was recognised as a distinct and new species in 1971.[2]


Amphilestes is known from various dental and mandibular remains. The dental formula of the mandible is 4:1:4:5. The premolars are symmetrical and the crowns look like tricusped molars, with the central cusp being largest in premolars and molars, though the size difference is less great in the premolars. George Gaylord Simpson (1928, p. 71)[3] noted that the teeth of Amphilestes are diagnosable in having “molar cusps high and slender, molar cingulum rising below the main cusp, molar enamel not pitted."[4]


  1. ^ Kermack, KA. 1988 British Mesozoic mammal sites. Special Papers in Palaeontology, 40:85-93.
  2. ^ Owen, R. 1871. Monograph of the Fossil Mammalia of the Mesozoic Formations. Palaeontographical Society.
  3. ^ G. G. Simpson. 1928. A Catalogue of the Mesozoic Mammalia in the Geological Department of the British Museum 1-215 [R. Benson/R. Benson/M. Uhen]
  4. ^ Butler, P.M. and Sigogneau-Russell, D. 2016. Diversity of triconodonts in the Middle Jurassic of Great Britain. Palaeontologia Polonica 67, 35–65. LSID pub: C4D90BB6-A001-4DDB-890E-2061B4793992