Temporal range: Late Triassic, 215Ma
Nesbitt et al., 2009
Nesbitt et al., 2009
Tawa (pron.:"Ta-WAH"; named after the Hopi word for the Puebloan sun god) is a genus of theropod dinosaur from the Late Triassic period. The discovery of Tawa alongside the relatives of Coelophysis and Herrerasaurus supports the hypothesis that the earliest dinosaurs arose in Gondwana during the Late Triassic period in what is now South America, and radiated around the globe from there. The specific name honours Ruth Hall, founder of the Ghost Ranch Museum of Paleontology.
Tawa was estimated to have been 2 m (6.5 ft) long, almost 2.5 feet at the hips. Other estimates suggest that Tawa weighed 40 kilograms (88 pounds) at most. The holotype, a juvenile individual, cataloged GR 241, consists of a mostly complete but disarticulated skull, forelimbs, the pectoral girdle, a partial vertebral column, hindlimbs, the pelvic girdle, ribs, and gastralia. The determination was made that this specimen is a juvenile based on the presence of an open braincase and unfused neurocentral sutures. An isolated femur GR 244 suggests that adults were at least 30% larger than the juvenile holotype. Tawa preserves characters that can be associated with different dinosaur taxa. It's skull morphology resembles that of coelophysoids and the illium approximates that of a herrerasaurid. Like the coelophysoids, Tawa has a kink in its upper jaws, between the maxilla and the premaxilla. With respect to limb proportion, the femur is very long compared to the tibia. A neck vertebrae adaptation in Tawa supports the hypothesis that cervical air sacs predate the origin of the taxon Neotheropoda and may be ancestral for Saurischians, and also links the dinosaurs with the evolution of birds. Compared to earlier dinosaurs like Herrerasaurus and Eoraptor, Tawa had a relatively slender build.
The following were assigned as paratypes: a nearly complete skeleton (GR 242); femora, pelvis and tail (GR 155); and cervical vertebrae (GR 243).
According to Nesbitt et al. (2009) Tawa can be distinguished based on the following features:
- the prootic bones meet on the ventral midline of the endocranial cavity
- the anterior tympanic recess is greatly enlarged on the anterior surface of the basioccipital and extends onto the prootic and the parabasisphenoid
- a deep recess is present on posterodorsal base of the paroccipital process
- a sharp ridge extending dorsoventrally on the middle of the posterior face of the basal tuber
- an incomplete ligamental sulcus is present on the posterior side of the femoral head
- a semicircular muscle scar/excavation is present on the posterior face of femoral head
- a small semicircular excavation on posterior margin of medial posterior condyle of proximal tibia
- a "step" is present on the ventral surface of the astragalus
- metatarsal I is similar in length to the other metatarsals
- the skull is elongate, three times longer than it is tall, and the antorbital fenestra in the skull is elongate, approximately 25% if the skull's length
- the premaxilla and maxilla are loosely articulated and the premaxillary tooth row ends anterior to external naris
- the anterior dentary teeth are enlarged
- the cervical epipophyses are narrow and project posterolaterally, there are no axial diapophyses, and the axial parapophyses are reduced or absent
- the pubic peduncle of ilium has two facets- one facing anteriorly and the other ventrally
- the distal tips of the pubic bones do not make contact
- the shape of the anterior trochanter is conical
Mortimer (2009) pointed out that Nesbitt et al.'s (2009) analysis excluded the above characters found by Tykoski (2005).
Fossils now attributed to Tawa were first discovered in 2004. Fossils of at least seven other individuals were also discovered at the site. One of these specimens, cataloged GR 242, is also nearly complete. All of these specimens are from the Hayden Quarry and date to between 215 to 213 million years old. Tawa was formally described in 2009 by a group of six American researchers led by Sterling J. Nesbitt of the American Museum of Natural History. At the time of publication in the journal Science, Nesbitt was a postdoctoral researcher at the University of Texas at Austin's Jackson School of Geosciences.
The type species is Tawa hallae, which was described in 2009 by Nesbitt et al., and considered more basal than Coelophysis, an early theropod from the Late Triassic. In 2009, Mortimer cautioned that Nesbitt et al.'s analysis was limited because it failed to consider all the characters of the relevant dinosaurs that the old analysis did (e.g. Guaibasaurus, Panphagia, "Dilophosaurus" sinensis, Dracovenator, Lophostropheus, etc.)Tawa was however found to be more advanced than the earliest theropod dinosaurs, Eoraptor and Herrerasaurus, and Staurikosaurus. Martinez et al. (2011) concluded that Tawa was the basalmost coelophysoid.
Sues et al. (2011) considered Tawa a derived early theropod. A cladistic analysis of Tawa and other early theropods indicate that the Coelophysoidea, a group of early dinosaurs, may be an artificial grouping because Tawa combines classic coelophysoid features with features which appear to be ancestral to the neotheropods. Tawa is believed to be the sister taxon of Neotheropoda, a group of carnivorous dinosaurs which bore only three functional digits on their feet.
Provenance and Occurrence
In 2004, a partial skeleton of Tawa was excavated, and in 2006 two nearly complete skeletons of Tawa and parts of six others were found close together in the Hayden Quarry at Ghost Ranchn in the lower portion of the Petrified Forest Member of the Chinle Formation in New Mexico. Tawa was discovered in gray/green siltstone from strata dating from the Norian stage of the Triassic period, approximately 215-213 million years ago. The discovery of Tawa alongside the relatives of Coelophysis and Herrerasaurus supports the hypothesis that the earliest dinosaurs arose in Gondwana during the Late Triassic period in what is now South America, and radiated around the globe from there.
Fauna and Habitat
Ghost Ranch was located close to the equator 200 million years ago, and had a warm, monsoon-like climate with heavy seasonal precipitation. Hayden Quarry, an excavation site at Ghost Ranch, has yielded a diverse collection of fossil material that included the first evidence of dinosaurs and less-advanced dinosauromorphs from the same time period. The discovery indicates that the two groups lived together during the early Triassic period 235 million years ago.Tawa's paleoenvironment included archosaurs, crocodylomorphs, rauisuchians, phytosaurs, ornithodirans including Silesaurus, and the dinosaurs Dromomeron, Chindesaurus, Eucoelophysis and possibly Coelophysis.(Irmis et al. 2007).
Based on their review of the early carnivorous dinosaur fauna from Ghost Ranch and the Ischigualasto Formation Nesbit et al. (2009) observed that each was descended from a separate lineage, and inferred that the "South American" protocontinent Gondwana was the ancestral range for basal dinosaurs. Nesbit et al. (2009) went on to note that dinosaurs left their ancestral range in Gondwana and 200 million years ago they dispersed across the adjoined continents of Pangea.
Nesbit et al. (2009) noted that repeated flooding events collected vertebrate bones, carcasses, and plant material from the landscape surface, possibly in hyperconcentrated flows, and deposited them at what is now Hayden Quarry. It was observed that these flooding events were separated by intervals where there was standing water and weakly-developed, poorly-drained (hydromorphic) soil formation. The Tawa specimens were very well preserved which suggests that they were buried extremely soon after dying.
- Maffly, Brian (December 10, 2009), "New Mexico find sheds light on early dinosaur dispersal", Salt Lake Tribune
- "New T. Rex Cousin Suggests Dinosaurs Arose in S. America", National Geographic, December 10, 2009
- Nesbitt, S. J.; Smith, N. D.; Irmis, R. B.; Turner, A. H.; Downs, A. & Norell, M. A. (2009), "A complete skeleton of a Late Triassic saurischian and the early evolution of dinosaurs", Science 326 (5959): 1530–1533, doi:10.1126/science.1180350, PMID 20007898.
- "TAWA". Hostgator.com. Retrieved 11 May 2013.
- Tykoski, R. S., 2005, Vertebrate paleontology in the Arizona Jurassic: In: Vertebrate Paleontology in Arizona, Edited by McCord, R.D., Mesa Southwest Museum Bulletin, no. 11, p. 72-93.
- DML: http://dml.cmnh.org/2009Dec/msg00122.html
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- Martinez, Sereno, Alcober, Columbi, Renne, Montanez and Currie, 2011. A basal dinosaur from the dawn of the dinosaur era in Southwestern Pangaea. Science. 331, 206-210.
- H.D. Sues, S. J. Nesbitt, D. S. Berman and A. C. Henrici. 2011. A late-surviving basal theropod dinosaur from the latest Triassic of North America. Proceedings of the Royal Society B 278:3459-3464
- Braginetz, Donna. "A new species of dinosauromorph (lower left) was among the mixed assemblage of dinosaurs and dinosauromorphs found at Hayden Quarry in Ghost Ranch, N.M.". American Museum of Natural History. Retrieved 31 March 2013.